Greece has tightened border controls, sending security forces to hold back migrants heading for the country by land and sea after Turkey warned that it would no longer prevent refugees from leaving.

Greece was the main gateway for hundreds of thousands of refugees, many fleeing fighting in Syria, who streamed out of neighbouring Turkey in 2015 and 2016 until a deal with the EU stemmed the flow.

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Greece sent more police to its northern border and increased maritime patrols around its islands on Friday, after Turkey said it would let out thousands of refugees stuck in the country by the 2016 deal.

A senior Turkish official said Turkey would no longer stop Syrian refugees from reaching Europe, in a response by Ankara to the killing of 33 Turkish soldiers in an air raid in Syria.

At one land crossing, police buses were seen denying access to dozens of people, including families with children, who had gathered in a no-man's-land between Pazarkule in Turkey and Kastanies in northeastern Greece.

"They will not enter the country. They are irregular migrants, we won't let them enter," a Greek government official told Reuters News Agency.

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Earlier on Friday, more than 60 migrants, most from Afghanistan, arrived on dinghies at the island of Lesbos, a Reuters witness said. Another boat carrying about 50 people reached the island of Samos, police said.

Greek Prime Minister Kyriakos Mitsotakis discussed the movement of migrants towards Greece with German Chancellor Angela Merkel.

"I want to be clear: No illegal entries into Greece will be tolerated. We are increasing our border security," Mitsotakis tweeted.

On Thursday, Greece had said it would tighten border controls to prevent coronavirus reaching its Aegean islands, where thousands of migrants are living in poor conditions.