A team spokesman said Reed was suffering from a sprained toe. Keeping him out of practice was believed to be a precautionary move, said a person with knowledge of the situation.

Reed, who missed time late in the season with a joint separation of his left shoulder, spent the offseason training primarily in Miami but did take part in the mandatory minicamp in June. He practiced without limitation.

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Reed last season earned Pro Bowl honors after recording 66 catches for 687 yards and six touchdowns. In 2015 he had 87 catches for 952 yards and 11 touchdowns.

While the regular season PUP list requires a player to miss six weeks before returning to practice, during training camp he can return to the field as soon as the medical staff clears him.

Meeting with reporters Wednesday, Coach Jay Gruden said third-round draft pick Fabian Moreau, a cornerback, and fourth-round pick Montae Nicholson, a safety, have progressed from offseason injuries and will be able to practice with the rest of the team when training camp opens. Both players recovered this offseason while their teammates practiced — Moreau from a torn pectoral muscle and Nicholson from shoulder surgery.

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Gruden said Wednesday that both players will be limited, meaning they initially will take part only in individual drills and not in any contact or team drills.

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Moreau, a 6-foot, 206-pound UCLA product, has a chance to compete for a key role in the secondary as a rookie, and it’s believed — based on his potential — that he could emerge as a starter in his second season. The 6-2, 212-pound Nicholson, who is from Michigan State, will likely contribute primarily on special teams early on.

Veteran safety DeAngelo Hall and outside linebacker Houston Bates both will open camp on the PUP list while still rehabbing from anterior cruciate ligament tears. Gruden said a couple of other players could start camp on the injured list as well, but those determinations had yet to be made.

The Redskins arrived at the Bon Secours training facility in Richmond Wednesday for a series of required conditioning tests. After meetings on Thursday morning, they will go through a walk-through, break for lunch, participate in classroom sessions and conduct their first practice in the afternoon.

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It marks the Redskins’ sixth summer in Richmond, and the fourth under Gruden. The coach and his players aim to build upon the success of the past two seasons, in which they had winning records back-to-back for the first time in 20 years.

Not since a stretch from 1989-92 have the Redskins recorded winning campaigns in three straight years, but Gruden expressed optimism that his squad is positioned to continue improving.

Washington returns quarterback Kirk Cousins and the bulk of the starters from an offense that ranked among the top five in the NFL in 2016. Team officials worked to upgrade a defense that struggled mightily last season, and after adding pieces in free agency and the draft, could field a starting lineup that features six to seven new faces.

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But banking on the strength of the team’s nucleus, and what he saw from the new additions during spring practices, Gruden feels confident in the roster.

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“We have great competition at every position — more than any time that I’ve been here. We want to sustain that competition day in and day out,” Gruden said. He later added: “I think the best part about our football team now is the core group of veterans that we have that have been here lead by example. You know the Trent Williamses of the world, the Kirk Cousinses, the Will Comptons. All these guys that have been here three and four years, they’re great leaders and they lead by example.

“And … then adding the free agents like Josh Norman, who is one of the hardest-working guys on our team, and watching how D.J. Swearinger works and Terrell McClain. … Adding some free agents that also love to work has been a great benefit for us,” Gruden said. “The character of this football team is the strongest it’s ever been, I believe. They get along great in the locker room, they work extremely hard on the practice field and hopefully that continues throughout the long process of training camp and preseason.”

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Cousins enters his third season as a starter, and for a second straight year will play on the franchise tag after team officials and his agent failed to reach an agreement on a long-term contract.

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But Gruden said he doesn’t expect the uncertain future to negatively affect Cousins’s play.

“He was fine last year; I thought he competed every day and did everything right,” Gruden said. “That’s the thing about Kirk, is you’re never worried about his preparation. It’s not going to vary from day in to day out. He’s not going to come in and be a different guy every day. He’s the same guy every day. A great competitor, wants to get better, understands he has a lot of room for improvement and he wants to be coached, and it’s fun to coach him.

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“The contract status will take care of itself eventually, hopefully,” Gruden said, “but right now, it is what it is and we’re happy to have him for 2017.”