The nearly 3-year-old Yemen war has killed more than 10,000 people, displaced 2 million and helped spawn a devastating cholera epidemic in the Arab world's poorest country. Amid its humanitarian crises, Yemen's culture and historical sites have also been affected.

The Awwam Temple links a region now on the front lines of the Saudi-led war against Houthi rebels to Arabia's pre-Islamic past.

Experts fear the temple, as well as other historic and cultural wonders across Yemen, beyond those acknowledged by international authorities, remains at risk as the country's stalemated war rages on.

"All the villages are historic in a way," said Anna Paolini, the director of UNESCO's regional office in Qatar that oversees Yemen and Gulf Arab nations. "They're still heritage of the country. It's sad to see what's happening."

Saudi-led air attacks have destroyed historic mud homes in Saada, the birthplace of the Houthi rebels. Air attacks have also hit the over 2,500-year-old Old City in Yemen's rebel-held capital of Sanaa, a UNESCO World Heritage Site for its intricately decorated, burnt-brick towers. Shelling and air attacks have struck museums and other sites in the country. In 2015, air attacks damaged part of the Great Marib Dam, near the Awwam Temple and built by the same civilization, according to UNESCO.

Just the shockwaves of an explosion in the distance can be enough to damage delicate structures. UNESCO has shared coordinates of some 50 historical sites with militaries involved in the fighting, to try to protect the sites, Paolini said, though many remain unguarded now in the chaos of the war.

"Although we've seen less collateral damage and targeting of heritage, it still happens," she said.

Foreign historians and archaeologists also fled the country over the fighting, halting work at sites like the Awwam Temple, also known as the Mahram Bilqis. Scholars believe the temple, with parts dating to the 7th century BC, served as a shrine to the god Almaqah.

The nearby city of Marib was the capital of the Saba dynasty, which ruled the area for centuries before the birth of Christ. The kingdom built the Great Marib Dam and controlled spice and incense trade routes. The legendary Queen of Sheba, said in the Bible and the Quran to have visited King Solomon in the 10th century BC, has been linked to Saba, though Ethiopians claim her as their own.

"She gave the king ... gold, large quantities of spices and precious stones. Never again were so many spices brought in as those the Queen of Sheba gave to King Solomon," the enigmatic Bible story goes. "King Solomon gave the Queen of Sheba all she desired and asked for ... Then she left and returned with her retinue to her own country." The Quran says Queen of Sheba was invited by King Solomon to abandon the worship of the sun and accept monotheism, to which she agreed.

The Sabaean dynasty later succumbed to challenges from other kingdoms, and the Great Dam gave away in the 6th century AD, which the Quran attributes to the Sabaean abandoning the worship of one God. Soon after, Islam quickly spread across the Arabian Peninsula and beyond.

Marib, which has grown into a city where many displaced by Yemen's war now live, and gunfire echoes through the day and night, has never been a particularly easy place in modern history. Famed American archaeologist and oilman Wendell Phillips began excavation of the Awwam Temple in 1951, but had to abandon all his equipment and flee raiding Bedouin tribesmen with his team.

Associated Press correspondent Wilton Wynn and his photographer wife Leila visited the temple in 1957 on a royal tour of the then-Kingdom of Yemen, apparently becoming the first Americans since Phillips to see the ruins. Wynn described Marib at the time as so remote "our plane was lost for more than a half hour trying to find it."

After the dam burst, "the city of Sheba was drowned and the kingdom never recovered," Wynn wrote in his AP dispatch. "Its population drifted away, leading to tribes moving to Syria, Mesopotamia and Palestine."

Today, a simple fence surrounds the temple, just off a heavily guarded road leading to a Saudi coalition airbase and an oil refinery. A watchman lives in a small makeshift shack against the fence with his children.

Foreign journalists and officials on a recent tour of Yemen organized by Saudi Arabia slipped through a large hole in the fence to marvel at the ancient South Arabian script inscriptions on stones inside. Some still bore what appeared to be red paint.

Eight limestone pillars towered over the site, though weathered, chipped and marred by the occasional bit of graffiti. A Yemeni militiaman with a Kalashnikov assault rifle over his shoulder quickly shimmied up between two pillars, looking down with a smile at was once one of ancient Yemen's holiest sites.