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One of Maymont’s otters has died. Pandora, a 17-year-old female North American river otter, died from natural causes Saturday, according to a statement from the historic estate and nature park in Richmond. She was one of two beloved otters on view in the Robins Nature & Visitor Center.

Pandora was acquired by Maymont in 1999 as a 6-week-old orphaned pup. It was perfect timing, as the nature center, featuring a large aquarium for river otters, was about to open.

Pandora, who died from kidney failure, was the first otter to join the center. Her mate, Neptune, is 19 years old and is still on view.

North American river otters typically live eight to nine years in the wild but have lived up to 21 years in captivity, according to the National Wildlife Federation. Pandora was known for her friendly nature, aquatic acrobatics and engaging audiences with her signature backflip.