Prosecutors in Peru have charged five men in the timber industry with the 2014 murders of four indigenous activists who had battled illegal logging in the Amazon jungle.

Two timber executives and three loggers have been charged with the shooting deaths of the activists, the prosecutor Otoniel Jara, who works in Peru’s remote Ucayali region, told the Associated Press on Wednesday.

Environmentalists say the case is unprecedented in Peru, where years of illegal logging and, on occasion, suspected attacks by those carrying it out have often been met with an ineffectual response from authorities.

“We hope that the legacy of the victims of this massacre can lead to justice,” said Tom Bewick, of Rainforest Foundation US, a group that funded efforts to bring the alleged killers to justice.

Bewick said he hoped the case would “set an example for other indigenous environmental defenders across the world”.

The indigenous group’s leader, Edwin Chota, along with Jorge Ríos Pérez, Leoncio Quinticima and Francisco Pinedo, were found dead on 1 September 2014.

Authorities say the men were killed with shotgun blasts in the Upper Tamaya-Saweto Asháninka indigenous territory along Peru’s border with Brazil.

The activists had defended the forests, traveling by canoe for three days to the regional capital city, Pucallpa, to file complaints and urge forestry officials to take action. They urged prosecutors to stop illegal logging, presenting photos and sketches they made of destruction they found.

Prosecutors say the five suspects could face up to 35 years in jail if convicted. The timber executives José Estrada and Hugo Soria are accused of ordering the killings, which were allegedly carried out by the loggers Eurico Mapes, Josimar Atachi and Segundo Atachi.

The Associated Press was not immediately able to reach the suspects or their attorneys for comment.

During the five-year investigation, the men have publicly denied the charges. They remain free and are believed to be living in the remote jungles of Peru.

Jara, the prosecutor, said prosecutors who were assigned to the case before him had abandoned it.

The prosecutor said the three loggers had been in the area where the bodies were found, while the two businessmen had lost revenue after the indigenous activists accused them of illegal logging.

On one occasion, Estrada, one of the timber executives, allegedly referred to Chota, telling witnesses: “I’ll pay whatever ... I want his head,” according to documents filed by prosecutors.

The bodies of Chota and Quinticima were found, while those of Ríos and Pinedo are still missing.

Global Witness has said 164 environmentalists were killed worldwide in 2018. About half were killed in Latin American countries including Brazil, Chile, Colombia and Guatemala.

Mauro Pío, an indigenous environmental leader who sought to protect Peru’s Amazon region, was shot dead in 2013. No suspects have been charged.

Relatives of the four Peruvian activists killed in 2014 said the jungle territory of their community remained vulnerable. But they welcomed the charges, hoping it signaled a shift toward more robust protections for indigenous groups.

“This is good,” said Asháninka Diana Ríos, daughter of Jorge Ríos, one of the murdered activists. “This is not going to be filed away and forgotten.”