A zoo in Denmark that provoked outrage after putting down a healthy giraffe has killed a family of four lions to make way for a new young male lion.

Copenhagen Zoo says it "had to euthanise" two cubs and their parents after it failed to re-home them.

The 16-year-old male and 14-year-old female were nearing the end of their natural lives in captivity, it added.

Malcolm Brabant reports.