1 / 19 Grand Prize/First Place: Action

Greg Lecoeur: "During the sardine migration along the Wild Coast of South Africa, millions of sardines are preyed upon by marine predators such as dolphins, marine birds, sharks, whales, penguins, sailfishes, and sea lions. The hunt begins with common dolphins that have developed special hunting techniques to create and drive bait balls to the surface. In recent years, probably due to overfishing and climate change, the annual sardine run has become more and more unpredictable. It took me two weeks to have the opportunity to witness and capture this marine predation."

Greg Lecoeur/2016 National Geographic Nature Photographer of the Year