This kind of change is what keeps Silicon Valley going. When personal computers displaced mainframe computers, it opened the door not just for Apple, but for companies making PC software for business, games and publishing. In the networking and Internet revolutions, venture capitalists invested in these new computing styles, and another generation of companies was born.

Over the last decade, smartphones, social networks and cloud computing have moved from feeding the growth of companies like Facebook and Twitter, leapfrogging to Uber, Airbnb and others that have used the phones, personal rating systems and powerful remote computers in the cloud to create their own new businesses.

Believe it or not, that stuff may be heading for the rearview mirror already. The tech industry’s new architecture is based not just on the giant public computing clouds of Google, Microsoft and Amazon, but also on their A.I. capabilities. These clouds create more efficient and supple use of computing resources, available for rent. Smaller clouds used in corporate systems are designed to connect to them.

The A.I. resources Ms. Greene is opening up at Google are remarkable. Google’s autocomplete feature that most of us use when doing a search can instantaneously touch 500 computers in several locations as it guesses what we are looking for. Services like Maps and Photos have over a billion users, sorting places and faces by computer. Gmail sifts through 1.4 petabytes of data, or roughly two billion books’ worth of information, every day.

Handling all that, plus tasks like language translation and speech recognition, Google has amassed a wealth of analysis technology that it can offer to customers. Urs Hölzle, Ms. Greene’s chief of technical infrastructure, predicts that the business of renting out machines and software will eventually surpass Google advertising. In 2015, ad profits were $16.4 billion.