Top American newspapers posted further declines in weekday circulation in the six-month period ended in March, with the exception of USA Today and The Wall Street Journal.

Apart from those two national dailies, which eked out gains of under 1 percent each, every other newspaper in the top 20 posted declines, according to figures released Monday by the Audit Bureau of Circulations.

USA Today, owned by Gannett, remained the top-selling paper in the country, with an average daily circulation of 2,284,219, up 0.3 percent.

The Wall Street Journal kept its No. 2 spot at 2,069,463, up 0.4 percent. The News Corporation, controlled by Rupert Murdoch, bought Dow Jones & Company, The Journal’s parent company, in December.