Leaders in Yemen’s southern separatist movement say they are “ready” for peace talks but will not abandon demands for an independent state, after their forces seized control of Aden city, the seat of the recognised government, in clashes with loyalist forces that left dozens dead.

Emirati-trained troops allied to the Southern Transitional Council last week captured army bases and the presidential palace in the strategic port city which serves as a temporary capital.

They clashed with Saudi-backed troops allied to the recognised president, Abedrabbo Mansour Hadi, ripping apart a Gulf coalition that was formed to fight the Houthi rebel group in the north of the country.

As Yemen teetered on the cusp of a second civil war, Abu Dhabi’s powerful crown prince met King Salman bin Abdelaziz and Crown Prince Mohammed bin Salman in Mecca on Monday, in an apparent effort to salvage the alliance.

Aidarus Zoubaidi, chief of the separatist Southern Transitional Council (STC), said late on Sunday that he was committed to a ceasefire in Aden and would join emergency talks in Riyadh.

However, Ahmed Bin Fareed, the movement’s representative in Europe, told The Independent that while they were keen to participate in peace talks they would not back down on calls that Yemen be divided, dashing hopes of a speedy resolution to the crisis.

Children suffering from acute malnutrition in Yemen Show all 17 1 /17 Children suffering from acute malnutrition in Yemen Children suffering from acute malnutrition in Yemen Doctors take some blood of Yemeni Yousef Abdullah Bakhit Ali, 13, suffering from severe acute malnutrition. With ongoing and unending conflict in Yemen, humanitarian situation continues to deteriorate across the country Unicef/Huwais Children suffering from acute malnutrition in Yemen A doctor weighs Yemeni baby Yahya Hamoud Ali Al Huzef, 9 months suffering from malnutrition Unicef/Huwais Children suffering from acute malnutrition in Yemen Ali Mohammed Ahmed Jamal,12 years old and suffering from severe acute malnutrition. He arrives with his family at a Unicef supported treatment centre in a hospital in Sanaa Unicef/Huwais Children suffering from acute malnutrition in Yemen A doctor measures the arm of Yemeni Ali Mohammed Ahmed Jamal, 12, who is suffering from malnutrition at a treatment centre in a hospital in Sanaa Unicef/Huwais Children suffering from acute malnutrition in Yemen There are over 400,000 severely malnourished children in need urgent lifesaving assistance in Yemen Unicef/Abdulhaleem Children suffering from acute malnutrition in Yemen Ali Mohammed Ahmed Jamal is weighed Unicef/Huwais Children suffering from acute malnutrition in Yemen A doctor measures the arm of baby Yahya Hamoud Ali Al Huzef Unicef/Huwais Children suffering from acute malnutrition in Yemen Unicef are currently working to reach 275,000 malnourished children with critical life-saving supplies and care for over 5 million people with safe and clean water to stop the spread of life-threatening diseases Unicef/Huwais Children suffering from acute malnutrition in Yemen Yahya Hamoud Ali Al Huzef with his family in his house in the outskirts of the capital Sanaa Unicef/Huwais Children suffering from acute malnutrition in Yemen The country is on the brink of famine and children's chances of survival are becoming slimmer by the day Unicef/Abdulhaleem Children suffering from acute malnutrition in Yemen Ali Mohammed Ahmed Jamal has his arm measured Unicef/Huwais Children suffering from acute malnutrition in Yemen Unicef are currently working to provide nearly 1 million children with vaccines and healthcare Unicef/Huwais Children suffering from acute malnutrition in Yemen Doctors take some blood of Yemeni Yousef Abdullah Bakhit Ali, 13 years old suffering from severe acute malnutrition at a treatment centre in a hospital in the capital Sanaa on November 2, 2018. (Release obtained) With ongoing and unending conflict in Yemen, humanitarian situation continues to deteriorate across the country. There are over 400,000 severely malnourished children in need urgent lifesaving assistance in Yemen. The country is on the brink of famine and childrenâ€™s chances of survival are becoming slimmer by the day. UNICEF are working with partners around-the clock to save children suffering from malnutrition and disease. We are currently working to reach: â€¢ 275,000 malnourished children with critical life-saving supplies and care â€¢ Over 5 million people with safe and clean water to stop the spread of life-threatening diseases â€¢ Nearly 1 million children with vaccines and healthcare â€¢ 9 million people with emergency cash assistance to help families buy basic commodities so they can survive Â© UNICEF/UN0253367/ Huwais Unicef/Huwais Children suffering from acute malnutrition in Yemen Unicef are working with partners around-the clock to save children suffering from malnutrition and disease Unicef/Abdulhaleem Children suffering from acute malnutrition in Yemen Unicef are currently working to provide 9 million people with emergency cash assistance to help families buy basic commodities so they can survive Unicef/Huwais Children suffering from acute malnutrition in Yemen Ali Mohammed Ahmed Jamal is suffering from malnutrition Unicef Children suffering from acute malnutrition in Yemen On 31 October 2018 in Yemen, the Al Thawra Hospital in Hodeidah where children are being treated for severe acute malnutrition. UNICEF Regional Director for Middle East and North Africa Geert Cappelaere visited Yemen from 29 October through 1 November 2018. As of 30 October 2018, over 11 million children â€“ 80 per cent of all children in the country â€“ require humanitarian assistance, due to the impact of the ongoing conflict. Worsening years of underdevelopment, attacks against civilians and civilian infrastructure and lack of salary payments for most civil servants have pushed basic services for children to the brink. Children face food shortages, disease, displacement and an acute lack of access to basic social services. One out of three children in Yemen is at risk of acute malnutrition. Â© UNICEF/UN0253574/Abdulhaleem Unicef/Abdulhaleem

“We accept to go to Riyadh for talks, we are still committed to the coalition, we respect President Hadi but we are sure he is controlled by members of the [Islamist] Islah Party,” Mr Fareed said on Tuesday.

“We will not back down from our demand of a state. No one will accept a unified Yemen. South Yemen was an independent state until very recently. We want a return to our country,” he added.

Thousands of southern separatists have received training and arms from the UAE as part of the effort to oust the Iran-backed Houthis who swept control of the country in 2014 ousting Mr Hadi.

The Houthi takeover of the country five years ago had prompted Saudi Arabia and its chief ally UAE to launch a bombing campaign to reinstate Mr Hadi, fearing encroachment of Iranian influence in the Gulf region.

However, fractures have started to show within the coalition, members of which have clashing agendas.

The southerners see a divided Yemen as the only future for the country and have called for a return to the borders of North and South Yemen, before the country was first unified in 1990.

The STC has also accused groups such as the Muslim Brotherhood-affiliated Islah, which backs Mr Hadi, of collaborating with the Houthis to scupper their aspirations. They point to a string of attacks on the STC’s forces, including a recent missile attack on a military parade near Aden which killed the group’s most senior commander, known by his nickname Aboul Yamama.

Mr Hadi’s government, meanwhile, has accused the UAE of backing a separatist “coup”.

To add to the confusion, Saudi-backed coalition warplanes hit separatist areas on Sunday and the coalition has vowed further attacks to dislodge southern fighters.

The UAE, whose crown prince rushed to talks with King Salman on Monday, meanwhile tried to downplay a rift with Riyadh, saying ties between the two Gulf allies remain strong and they would “adamantly confront” any powers that threaten regional security.

“The UAE and Saudi Arabia call on conflicting Yemeni parties to prioritise dialogue and reason for the interest of Yemen,” state news agency WAM quoted UAE crown prince Sheikh Mohammed bin Zayed al-Nahyan as saying after the meeting.

However, he stopped short of asking the southern fighters to cede control of the Yemeni port in favour of President Hadi’s government, a solution sought by Saudi Arabia.

Soldiers lie on the ground after a missile attack on a military parade during a graduation ceremony for newly recruited troopers in Aden, Yemen (Reuters)

The situation in Aden has calmed and civilian flights returned on Tuesday. However, the International Committee of the Red Cross warned that hospitals were in desperate need of supplies after 72 hours of heavy fighting left more than 70 dead.

It said many of the wounded were trapped during the clashes and could not reach health facilities, with some dying as a result. It added that some 200,000 were left without access to water.

There has been no movement of the separatist forces from the bases they seized.

Mr Fareed said that the eventual separation of Yemen was “inevitable” and there would be no peace until it happened.