JACKSONVILLE, Fla. (AP) — Jacksonville Jaguars defensive ends Yannick Ngakoue and Dante Fowler had to be separated repeatedly following practice Sunday.

Ngakoue and Fowler exchanged words while walking off the practice field, prompting the first skirmish. Calais Campbell grabbed Ngakoue briefly as Abry Jones and Lerentee McCray held back Fowler. Ngakoue spun away from Campbell and sprinted toward Fowler, but Jones got between them.

Ngakoue and Fowler got into it again outside the practice-field gates and had to be pulled apart by teammates, coaches and security personnel. General manager Dave Caldwell could be seen leading Ngakoue away by the wrist.

The altercation came on the last of 11 consecutive practices in full pads and in sweltering heat and humidity. It also came a few minutes after a scuffle near the end of practice, when Fowler appeared to shove tight end James O’Shaughnessy out of bounds following a reception. Teammates rushed to break it up.