WASHINGTON (AP) — Two Florida businessmen tied to President Donald Trump's lawyer and the Ukraine investigation were charged Thursday with federal campaign finance violations. The charges relate to a $325,000 donation to a group supporting Trump's reelection.

Lev Parnas and Igor Fruman, associates of Rudy Giuliani, were arrested Wednesday trying to board an international flight with one-way tickets at Dulles International Airport in Virginia, according to Geoffrey Berman, the U.S. attorney in Manhattan. No destination was disclosed.

Parnas and Fruman were arrested on a four-count indictment that includes charges of conspiracy, making false statements to the Federal Election Commission and falsification of records. The men had key roles in Giuliani's efforts to launch a Ukrainian corruption investigation against Democratic presidential contender Joe Biden and his son Hunter.

The indictments mark the first criminal charges related to the Ukraine controversy. While they do not suggest wrongdoing by the president, they are likely to add fuel to the House impeachment inquiry, raising additional questions about whether those close to Trump and Giuliani sought to use their influence to affect U.S. foreign policy decisions.

Trump has dismissed the impeachment inquiry as baseless and politically motivated.

Records show that Parnas and Fruman used wire transfers from a corporate entity to make the $325,000 donation to the America First Action committee in 2018. But wire transfer records that became public through a lawsuit show that the corporate entity reported as making the transaction was not the source of the money.

Prosecutors also allege that Parnas urged a congressman to seek the ouster of the U.S. ambassador to Ukraine, at the behest of Ukrainian government officials. That happened about the same time that Parnas and Fruman committed to raising more than $20,000 for the politician.

The congressman wasn't identified in court papers, but the donations match campaign finance reports for former Rep. Pete Sessions, a Texas Republican who lost his re-election bid in November. In May 2018, Parnas posted a photo of himself and his business partner David Correia with Sessions in his Capitol Hill office, with the caption "Hard at work !!"

John Dowd, an attorney for Parnas and Fruman, hung up on an Associated Press reporter seeking comment. Giuliani said he couldn't comment and that he didn't represent the men in campaign finance matters.

The men were arrested around 6 p.m. Wednesday and booked at a local jail in Alexandria, Virginia. A court appearance Thursday was delayed as lawyers tried to work out a bail package. Kevin Downing, the lawyer who represented former Trump campaign chairman Paul Manafort on charges that he hid millions of dollars that he earned in Ukraine advising politicians there, was representing the men for their initial appearance and declined to comment.

Correia and Andrew Kukushkin, a Ukrainian-born U.S. citizen, were also charged in the case.

Attorney General William Barr had been briefed on the investigation soon after he was confirmed in February, was updated in recent weeks and was made aware Wednesday night that the men were being arrested, a person familiar with the matter told The Associated Press. The person spoke on condition of anonymity to discuss an ongoing investigation.

The indictment says Parnas and Fruman "sought to advance their personal financial interests and the political interests of at least one Ukrainian government official with whom they were working" and took steps to conceal it from third parties, including creditors. They created a limited liability corporation, Global Energy Producers, and "intentionally caused certain large contributions to be reported in the name of GEP instead of in their own names."

Prosecutors charge that the two men falsely claimed the contributions came from GEP, which was described as a liquefied natural gas business. At that point, the company had no income or significant assets, the indictment said.

Prosecutors allege that Parnas and Fruman conspired to make illegal contributions to try to skirt the limit on federal campaign contributions. The men are also accused of making contributions to candidates for state and federal office, joint fundraising committees and independent expenditure committees in the names of other people.

The commitment to raise more than $20,000 for the congressman was made in May and June 2018. The lawmaker had also received about $3 million in independent expenditures from a super political action committee that Parnas and Fruman had been funding. A super PAC can raise and spend unlimited amounts of money in support of a candidate but isn't allowed to directly coordinate with the official campaign.

As a result of the donations, Parnas and Fruman had meetings with the congressman and Parnas lobbied him to advocate for removing the ambassador to Ukraine, Berman said. Trump referred to Ambassador Marie Yovanovitch, who was indeed recalled to the U.S., as "bad news" in his July phone call with Ukrainian President Volodymyr Zelenskiy.

Berman said his office "will not hesitate to investigate and prosecute those who engage in criminal conduct that draws into question the integrity of our political process." His office had brought unrelated charges against the president's former legal "fixer" Michael Cohen last year.

The indictment also charges that Kukushkin conspired with the three other defendants to make political contributions, funded by a foreign national, to politicians seeking state and federal office "to gain influence with candidates as to policies that would benefit a future business venture."

An unnamed foreigner wired $500,000 from a bank account overseas through New York to the defendants for contributions to two candidates for state office in Nevada, the indictment alleges. Foreigners are not permitted to contribute to U.S. elections.

The indictment accuses the four men of also participating in a scheme to acquire retail marijuana licenses through donations to local and federal politicians in New York, Nevada and other states.

The big donation to the Trump-allied PAC in May 2018 was part of a flurry of political spending tied to Parnas and Fruman, with at least $478,000 in donations flowing to GOP campaigns and PACs in little more than two months.

The money enabled the relatively unknown entrepreneurs to quickly gain access to the highest levels of the Republican Party, including meetings with Trump at the White House and Mar-a-Lago in Florida.

America First Action said the $325,000 contribution would remain in a separate account while the court cases play out. A spokeswoman, Kelly Sadler, said the committee will "scrupulously comply with the law."

The AP reported last week that Parnas and Fruman helped arrange a January meeting in New York between Ukraine's former top prosecutor, Yuri Lutsenko, and Giuliani, as well as other meetings with top government officials.

Giuliani's efforts to launch a Ukrainian corruption investigation were echoed by Trump in the July 25 call with Zelenskiy. That conversation is now at the heart of the impeachment inquiry .

House Democrats subpoenaed Parnas and Fruman on Thursday for documents they have refused to produce to three House committees. The panels have also subpoenaed Giuliani.

A whistleblower complaint by an unnamed intelligence official makes reference to "associates" of Giuliani in Ukraine who were attempting to make contact with Zelenskiy's team, though it's not clear that refers to Parnas and Fruman. That could put the two men squarely in the middle of the investigation into Giuliani's activities.

Neumeister reported from New York City. Associated Press writers Colleen Long and Brian Slodysko in Washington, Larry Neumeister and Jonathan Lemire in New York and Matthew Barakat in Alexandria, Virginia contributed to this report.