LSU coach Les Miles indefinitely suspended starting quarterback Jordan Jefferson and reserve linebacker Josh Johns this morning shortly after Baton Rouge, La., police issued arrest warrants for the pair.

The two are among four Tigers players accused in a bar fight that sent four people to hospitals on Aug. 19, including one person who had three fractured vertebrae.

"As sad as this incident is,"

released by the Tigers, "it's important that we learn from it and take away a valuable lesson. This has weighed on all of us. It's time for us to come together as a team and focus on what we are here to do."

The two players were arrested on second-degree battery accusations, which are felonies. The

bail was set at $5,000 a piece for the two. Both players were bonding out about 9:40 a.m. Portland time.

The arrests and suspensions substantially darken the cloud over No. 4 LSU a week before its season opener against No. 3 Oregon on Sept. 3 in Arlington, Texas. Jefferson, Johns and two other Tigers teammates - defensive tackle Chris Davenport and wide receiver Jarvis Landry - are accused of participating in the bar fight that left four men injured. On Thursday the Tigers announced

for violation of an NCAA rule in an unrelated incident. He is expected to miss the Oregon game and possibly others. Shepard had been interviewed about the NCAA’s ongoing investigation into the relationship that scouting services provider and purported street agent Willie Lyles has had with a number of college football programs, including both LSU and Oregon.

On Wednesday Baton Rouge police took a DNA swab from Jefferson and

. Police would not say why they wanted the shoes but a source close to the investigation said police were looking to match a shoe to a shoeprint on one of the victims' shirts. A police report says a 19-year-old woman named Victoria Long said she witnessed several LSU players beating one of the alleged victims and saw Jefferson kick that man in the face.

Two witnesses who work at the bar told The Associated Press that the players did not throw the first punch, but added that once the melee began, it was difficult to see who hit whom, and could not verify the extent to which Jefferson and Johns were involved.

Jordan Neldare, the door manager of the bar, who said he witnessed the fight, said he noticed that Jefferson removed himself from the brawl before it was over, and was standing alone near the bar’s entrance, looking upset.

As quarterback in 2010, Jefferson was 118 of 209 passing for 1,411 yards with a 56.5 completion percentage and a 114.7 passer rating.

His primary backup and likely replacement, Jarrett Lee, was 54 of 89 for 573 yards, 2 TDs 1 interception with a 60.7 completion rating and 119.9 passer rating in 2010. On his career, Lee has thrown for 2,643 yards, 17 touchdowns and 18 interceptions.

The Tigers' other backup quarterback, Zach Mettenberger, has not seen game action at LSU, transferring to Baton Rouge this spring after a year at Butler (Kansas) Community College. Before that he was at Georgia, but did not ever take the field for the Bulldogs.

Under team orders, LSU players have not discussed the fight publicly since it happened.

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- This report was assembled with feeds from The Associated Press, The New Orleans Times Picayune, select beat reporters via Twitter and the LSU Tigers official site.