Here are five storylines to follow when the teams meet at FedEx Field.

1. Revamped offensive line: The loss of eighth-year center Kory Lichtensteiger and injury to left guard Shawn Lauvao will test the coaching ability of offensive line coach Bill Callahan and the readiness of two young players: Spencer Long, who’ll take over at center, and Arie Kouandjio, who’s expected to get his first NFL start in Lauvao’s spot. The upshot is a line with staggering size, with no player under 6-feet-5. Long has 13 NFL starts at left guard, but Cousins says he’s confident in his ability to make the calls at center and protect up front. At 6-5 and 324 pounds, Long is three inches taller and 32 pounds heavier than Lichtensteiger. The Redskins also signed nine-year NFL veteran center John Sullivan as insurance.

2. Red-zone woes: The Redskins offense ranks fifth in total yards (406.3 yards per game), but the statistic amounts to little given the struggles inside the 20-yard-line. They’ve scored touchdowns just 21 percent of the time in the red zone, which ranks 31st in the NFL. To date, there’s no single reason for the red-zone struggles. The offense has fallen short in different ways, including poor throws, predictable play-calling and failed runs. Kicker Dustin Hopkins has been the saving grace, particularly in last week’s 29-27 victory at the New York Giants, connecting on all five of his attempts to earn NFC special teams player of the week honors. But it’s on offensive coordinator Sean McVay and Cousins, in particular, to start converting their red-zone chances.

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3. Sticking with the running game: After ignoring their running game in Weeks 1 and 2, the Redskins followed a more balanced offensive script against the Giants and were rewarded in many ways. They gained a modest 90 yards on the ground in the seesaw battle, with second-year back Matt Jones carrying 17 times for 65 yards. But for the first time this season, they won time of possession, with the offense staying on the field for 33:26 and, in turn, helping the defense stay fresh. The Redskins won’t win many games if Cousins throws 75 percent of the time. To be effective, they have to give defenses more to worry about than Cousins’s arm and his corps of receivers.

4. Rookie CB Kendall Fuller: The Redskins opened the season with 11 defensive backs. Heading into Week 4, they sorely need the depth. Three cornerbacks missed all or a portion of practice this week: Bashaud Breeland (ankle), Josh Norman, fending off what Coach Jay Gruden hopes is a flu bug, and Dashaun Phillips (hamstring). Safety DeAngelo Hall is out for the season with a torn ACL. The re-shuffling in the backfield means that Fuller, a third-round pick from Virginia Tech, should finally get in a regular-season game Sunday after being edged by Phillips for the nod in training camp. Now fully recovered from last fall’s knee surgery, Fuller is “more than ready,” Gruden declared.