CINCINNATI -- At the close of their first day of training camp, the Cincinnati Bengals made a series of roster moves that included signing two players, releasing another and placing a fourth on the active physically unable to perform list.

Pro Bowl offensive tackle Andrew Whitworth was the PUP list addition, the result of a calf injury. Like the rest of the PUP list players, he counts against the team's roster limit and can be activated to practice as soon as he's cleared medically.

Whitworth wasn't the only one who sat out Thursday's practice. Tight end Jermaine Gresham (back), quarterback AJ McCarron (shoulder), offensive lineman Mike Pollak (knee), receivers Marvin Jones (ankle) and Ryan Whalen (hamstring), linebacker Marquis Flowers (hamstring), and defensive tackles Devon Still (back) and Zach Minter (back) also didn't practice. Each had been placed on the active PUP or active non-football injury lists before training camp began.

While the Bengals were making those injury designations, they also made alterations to their roster. Backup punter T.J. Conley, who was signed earlier this offseason to primarily give the Bengals an extra practice body to keep starting punter Kevin Huber's leg fresh, was released. In a corresponding move, they added two receivers, Jeremy Johnson and Conner Vernon, who was acquired from Cleveland after clearing waivers.

Johnson is a rookie from SMU who signed with the Patriots as an undrafted free agent last month. He was released by New England on July 17.

Vernon entered the NFL in 2013, signing with the Raiders as an undrafted player. He had four catches for 42 yards in four preseason games last year before getting cut the final weekend of camp. He was signed by the Browns on Dec. 26, and remained part of their offseason roster.

With Jones and Whalen out for now, the Bengals needed a pair of receivers to take their places until they return. Although Johnson and Vernon will try to make the 53-man roster, they were added primarily for camp depth concerns.