Image copyright Declan McCarthy Image caption Passengers queue to get on to a South Western Railway train

Workers for three rail operators are continuing their 48-hour strike action.

Rail, Maritime and Transport (RMT) union members working for Southern, Greater Anglia and South Western Railway walked out on Wednesday.

The union is in dispute over driver-only operated (DOO) trains, also known as driver-controlled operated (DCO).

RMT staff on Merseyrail and Northern also staged a 24-hour walkout as part of Wednesday's strike action, which saw delays and disruption for passengers.

In a separate dispute, union Aslef said its drivers had voted to accept a deal to end their dispute with Southern Rail over driver-operated trains.

Members of the RMT union are mostly conductors.

Greater Anglia said it was running a full service during the industrial action, using other trained staff in place of conductors.

Some trains between Marks Tey and Sudbury had earlier been cancelled, but the service was running normally by about 16:00 GMT, a Greater Anglia spokeswoman said.

Southern said services on most of its routes were operating normally, although there were some alterations. However, there were delays in and out of Brighton Station in the morning due to an "obstruction" on the track, it said.

There is also disruption between Littlehampton and Southampton due to a points failure.

RMT's dispute with Southern's parent company, Govia Thameslink (GTR), over DOO trains has seen a spate of strikes since April 2016 amid concerns over safety and job losses.

Changes were introduced on Southern in January to make conductors "on-board supervisors" and pass responsibility for closing doors to drivers.

Industrial action by RMT members has recently spread to other routes across England.

Image copyright @PCSEastern Image caption RMT picketers on the second day of strike action at Norwich station

Richard Dean, Greater Anglia Train service delivery director, said: "The Office of Rail and Road is satisfied with our arrangements for training additional staff as conductors.

"We will never compromise the safety of our customers.

"We are running a full service again on Thursday."

South Western Railway, which only took over the franchise from South West Trains in August, is running more than 60% of services, and has published a contingency timetable, including replacement buses on some routes.

Image copyright PA Image caption Commuters ride a crowded South Western Railway train on the Portsmouth to London line on Wednesday

On Wednesday, RMT general secretary Mick Cash said: "The only way that Greater Anglia can be running these services is through taking serious risks with public safety just as they did during the last phase of strike action.

"Rail companies are training up rail staff who have previously had no rail operational experience to stand in as highly trained guards."

A spokesman for the Department for Transport said on Wednesday: "This dispute is not about jobs or safety - employees have been guaranteed jobs and salaries. In fact at Southern Rail, where these changes have already been introduced, there are now more staff on trains.

"The independent rail regulator has said driver-controlled trains, which have been used in this country for more than 30 years, are safe."

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