Greenpeace divers were inspecting Tubbataha's reefs for damage

Their flagship Rainbow Warrior II ran aground at Tubbataha Reef Marine Park, in the Sulu Sea, 650km (400 miles) south-east of Manila.

Park officials said almost 100 sq m (1,076 sq ft) of reef had been damaged.

Greenpeace agreed to pay the fine, but blamed the accident on outdated maps provided by the Philippines government.

"The chart indicated we were a mile and a half" from the coral reef when the ship ran aground, regional Greenpeace official Red Constantino told AFP news agency.

"This accident could have been avoided if the chart was accurate," he said, adding, however, that Greenpeace felt "responsible" for the damage.

'Immediate action'

The accident happened while the Rainbow Warrior was on a four-month tour of the Asia-Pacific region to promote environmentally-friendly energy sources.

Greenpeace divers were at the Tubbataha park, off the coast of Palawan island, to inspect the effect of global warming on the coral reef.

The Rainbow Warrior escaped serious damage

The Rainbow Warrior II escaped serious damage and was towed into deeper water by its own rubber boats.

Tubbataha park manager, Angelique Songco, praised the work Greenpeace was doing to protect the environment.

"We also appreciate the immediate action they took to get the full assessment of the damage," she said.