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Neptune, a male North American river otter at Maymont who lost his longtime female companion just more than a month ago, has died, too.

The otter, who was 19, died Friday from natural causes, Maymont said Wednesday.

The day before his death, Neptune was swimming, playing and behaving normally at the Robins Nature and Visitor Center, Maymont said in a news release.

Neptune’s death follows that of Pandora, a 17-year-old female otter who died of kidney failure Sept. 18.

North American river otters typically live eight to nine years in the wild but have lived up to 21 years in captivity, according to the National Wildlife Federation. Maymont attributed the pair’s long lives to constant attention and medical care from the nature center’s staff.

Neptune came to Maymont in 1999, when he was 2 years old. He previously had been hand-raised at a nature attraction in Silver Springs, Fla.

Pandora also arrived at Maymont in 1999.

The otters immediately bonded and became two of the most popular animals at the nature center, Maymont said.