“What I’m doing right now is madly preparing for the winter study, because of course it could happen,” Vucetich, an ecologist at Michigan Technological University, said Thursday. “It just might not happen.”

That’s because Isle Royale is a national park that is inaccessible to Vucetich and his colleagues because of the shutdown, which has also hobbled other scientific work. The study — which mainly consists of counting moose and the wolves that prey on them and observing the animals' interactions — depends on winter conditions and on a helicopter contract and pilot that cannot easily be extended after the planned end date, he said.

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A shortened study period would amount to lower quality data, Vucetich said. “If we are indeed unable to count wolves and moose this year,” he added, “that will be a missing spot in the database forever.”

The shutdown might also interrupt a National Park Service-led project to relocate as many as 30 wolves to Isle Royale, a 200-square-mile ecosystem where wolf numbers have dropped from a peak of 50 in the early 1980s to just one highly inbred pair. The first four wolves were moved and released in the fall, and one later died; the Park Service planned to release others this month.

The plan is aimed at restoring a predator-prey relationship that has long ebbed and flowed but is now severely out of balance: As of last year, there were 1,475 moose on Isle Royale and just the two wolves to control them. The moose population is growing by 15 percent a year, Vucetich said, and they are devastating the wooded island’s vegetation.

Moose came to Isle Royale in the early 20th century, probably by swimming to it, and wolves arrived in the late 1940s by walking across an ice bridge from Canada. After its peak, the wolf population plummeted because of disease, and the moose population mushroomed. The moose population fell and rose again, while the wolves have struggled with inbreeding. The ice bridges that mainland wolves might use to move to the island, meanwhile, are forming less frequently because of a warming climate, the Park Service says.

Scientists have been closely watching these dynamics for six decades. What does one abbreviated or missed survey matter in such a long-term effort? Maybe very little, Vucetich said, but maybe a lot.

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Vucetich compared the study to watching a movie: If you miss a few minutes to get popcorn, you might come back and find it easy to follow. “But if you miss the wrong two minutes, you’ll come back and say, ‘I have no idea what’s going on,’” he said, because you’ve missed a key plot twist.

“Two of the most important things we’ve learned on Isle Royale in 60 years are those kind of singular events,” he said.

The first was the collapse of the wolf population in the early 1980s, which was caused by the inadvertent introduction by humans of canine parvovirus, a disease that can also kill domestic dogs. The second was the migration of a male wolf that left Canada in 1997, traversed the iced-over Lake Superior and “bolstered pretty importantly the genetic fitness of the wolf population,” Vucetich said.

“Every winter, we try to get an impression of what it is that the wolves are doing. And the more time you have, the better you can do it,” he said.

This year’s study may be particularly important because of the introduced wolves, all of which are female. Because of the change in the wolf-moose ratio, the team also planned to put radio collars on some moose for the first time in decades, Vucetich said. Doing so is logistically complicated — involving two additional helicopters and more staff than the half-dozen people who are typically involved — so “there is a non-trivial risk that that effort to collar moose would be just scrubbed this year,” Vucetich said.

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The Park Service has plans to relocate additional wolves from Ontario to Isle Royale this month. Alexandra Picavet, communications chief for the National Park Service Midwest Region, said in a statement that because the effort requires Park Service employees and facilities, it is on hold “while there is a lapse in funding for the National Park Service.”

If it doesn’t happen this winter, it will be pushed back to the fall, Vucetich said. Spring and summer are off the table because, among other reasons, wolves have pups in the spring, and Isle Royale is open to visitors in the spring and summer.

Another few months without new wolves might seem minor in the big picture, he said.

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But “with no delays, we’re already looking at approximately a decade-long period with negligible wolf predation as an influence,” Vucetich said. “Every year that you wait, you increase damage you can’t undo.”