The four alleged poachers who survived were arrested and will appear in court “in due course,” the statement said. They will remain in custody until Friday, pending bail, reported Times Live, a South African news site. Authorities have launched an investigation into the alleged poacher’s death and accomplices.

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Park officials did not immediately respond to a request for comment from The Washington Post.

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Police Brig. Leonard Hlathi told Times Live that the elephant attacked “suddenly,” killing the man.

“His accomplices claimed to have carried his body to the road so that passersby could find it in the morning,” Hlathi said. “They then vanished from the park. ”

The suspected poachers called the dead man’s family, which then called Don English, regional ranger for Skukuza — the most popular game reserve in the country. English assured the family that he would “do everything possible” to recover the body, according to the park statement.

A search party, including rangers on foot and a crew with the South African National Parks air wing, scoured the area Wednesday but did not find the man. They resumed their efforts Thursday and found the man’s remains in the Crocodile Bridge area of the park, near the Crocodile River.

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The area, according to the Kruger National Park website, is known for its high concentration of lion prides. It also houses a large percentage of the park’s rhino population.

All that the lions had left behind of the man was his skull and a pair of pants, authorities said.

In the park statement, Phillips commended the search party for helping the family find closure. He also offered the family his condolences.

"It is very sad to see the daughters of the deceased mourning the loss of their father, and worse still, only being able to recover very little of his remains,” Phillips said.

When the other suspected poachers were arrested, authorities seized two .375 hunting rifles and ammunition. The men were charged with possessing firearms and ammunition without a license, conspiracy to poach and trespassing, Times Live reported.

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South Africa is home to the continent’s largest rhino population — about 20,000 of the 25,000 living in Africa. It holds nearly 80 percent of the world’s rhino population.

Although poaching numbers have been steadily declining in recent years, more than 8,000 rhinos have been killed in the past decade. Poaching numbers were most severe between 2013 and 2017, when more than 1,000 rhinos were killed in South Africa annually, according to statistics from Save the Rhino. Those numbers dipped below 1,000 in 2018, with 769 total poachings.

Save the Rhino reports that half of all poachings occur in Kruger National Park.