The university began its own investigation, and has interviewed members of the club and met with its leadership. “We can’t go into specifics or determine exactly when it will end,” said Martin Mbugua, the university’s spokesman.

Amid complaints from members, some describing the emails as evidence of a climate that is hostile toward women, the club’s graduate board of governors conducted a survey that invited respondents to strongly agree, agree, disagree or strongly disagree to statements such as “Given the current culture at Tiger Inn, it would NOT surprise me to hear that a sexual assault occurred at the club.” The survey did not refer to any specific episode.

Last week members were notified of the results. “After carefully listening to all sides — and to you,” the board wrote, “it is clear to us that the actions taken by Adam Krop and Drew Hoffenberg in the second week of October were offensive, disrespectful and in direct violation of our core values. This afternoon we asked Adam and Drew to step down as vice president and treasurer.” The letter announced additional measures, including “a slate of officers that is more balanced by gender” and “a safe process for members to report incidents or concerns.”

Neither of the two former officers, who are both seniors, responded to requests for comment Monday.

In an email, the head of the graduate board, Hap Cooper, said: “The survey found that a large majority of T.I. members are comfortable within their own skin at the club. It surfaced, however, a number of specific events, behaviors and ceremonies that demonstrate gender bias, and as such, must be modified or eliminated.”

The survey comes as the treatment of female students has gained urgency on campuses around the country. A report in Rolling Stone about a fraternity gang rape has roiled the University of Virginia, which suspended all fraternity and sorority activity through the end of the year. On Monday, Wesleyan University announced a ban on social activity for a fraternity where sexual assaults were reported. And the federal Education Department is investigating sexual violence cases at 90 institutions of higher learning.