The San Francisco 49ers announced on Thursday that they have signed linebacker Philip Wheeler to a one-year deal. The team also released wide receiver Trindon Holliday and waived guard Fouimalo Fonoti and wide receiver Lance Lewis.

Wheeler (6-2, 245) was originally drafted in the third round (93rd overall) of the 2008 NFL Draft by the Indianapolis Colts. Throughout his seven-year career, he has spent time with the Colts (2008-11), Oakland Raiders (2012) and Miami Dolphins (2013-14). Wheeler has played in 108 games (60 starts) and has registered 455 tackles, 5.5 sacks, four forced fumbles and two fumble recoveries. In 2013 with the Dolphins, he led the team with a career-high 118 tackles.