In Yesterday Origins, the main protagonists John Yesterday and his partner Pauline, immortals who own an antiques store in Paris, travel across the globe through multiple periods. The story takes place across the current and medieval eras, through a series of flashbacks. At the beginning of the game, a young John is imprisoned by the Spanish Inquisition for witchcraft, just prior to the sequence of events that will change his life/lives forever. Although not yet able to resurrect at that point of his life, he has already attracted the attention and fallen foul of Church authorities due to his unusual ability to understand and speak every language he encounters.

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