Level 1:

-Block: Periodically reduces the damage received from Hero basic attacks by 50%. Stores up to 2 charges.

-Regeneration Master: Increases health regeneration by 4 per second. Every 3 regeneration globes gathered increases this bonus by 4.

-Searing Flame: Auto attacks deal 25% extra damage while channeling Roaring Flame.

-Tuning Fork: Increases the duration of Modulation by 2 seconds and increases the Sound granted by basic attacks against Atramedes while Modulation is active to 3.

Level 4:

-Sighted, But Blind: Enemy heroes that grant Atramedes Sound by attacking him while Modulation is active are slowed by 10% for 2 seconds.

-Death Waits In The Darkness: Damage dealt by Atramedes increases up to 20% the smaller his field of vision from his most recent Sonar Pulse is.

-Superiority: Reduces damage taken from Minions, Structures, and Mercenaries by 50%.

-Deafen: Enemies near Atramedes when he activates Sonic Boom are stunned for 1 second.

Level 7:

-Loud And Thunderous: Increases the vision radius granted by Sonar Pulse by 30%

-You Cannot Run: Increases the speed bonus from Sonic Boom to 25% and reduces the Sound consumed per second to 3.

-Mr. Magma-Goo: Searing Flame sticks to enemies, burning them for an additional 30% damage over 3 seconds.

-Battle Momentum: Basic attacks reduce ability cooldowns by 0.5 seconds.

Level 10:

(R1) Resonating Roar: After a 1.5 second delay Atramedes unleashes a terrible roar that stuns enemy heroes for 2 seconds and deals massive damage to nearby structures. Consumes 30 Sound .

(R2) Hide & Seek: Causes all enemy heroes to glow (that distinct 'glow-n-the-dark' green color) until they have traveled X distance. Glowing enemies are visible in the fog of war but not on the minimap.

Level 13:

-No Need For Eyes: Every second Sonar Pulse becomes empowered, causing cloaked enemy heroes revealed to Atramedes by Sonar Pulse to be knocked out of stealth.

-Little Hearts Beating: Enemy Heroes below 50% health are visible for 3 seconds after leaving Atramedes' field of vision.

-In Synch: Modulation also shields a nearby ally.

-Spell Shield: Upon taking ability damage, reduce that damage and further ability damage by 50% for 2 seconds. Can only trigger once every 30 seconds.

Level 16:

-You Cannot Hide: Any enemy hero that causes Atramedes to generate Sound burns for X fire damage over two seconds.

-Malkoriak's Mixture: Reduces the Sound per second consumed by Roaring Flame by 5.

-Foul Stench (Active): Activate to smell the area, revealing the path of enemy heroes who have been in the area in the last 10 seconds. Path is visible for 5 seconds. 60 second cooldown.

-Clumsy Footsteps: Enemy Heroes, mercenaries and minions are slowed by 20% for 1 second when they pass through the threshold of Sonar Pulse.

Level 20:

-Cacophony: Removes the casting time from Resonating Roar and Resonating Roar is cast when Atramedes dies.

-Ready Or Not: Reduces the cooldown of Hide & Seek by 30% and enemy heroes affected by Hide & Seek leave a glowing tracer in their wake.

-Miserable Existence: Reduces time it takes for Atramedes to revive by 5 seconds for every hero that was visible to him when he died.