BENGHAZI, Libya — Tim Hetherington, a conflict photographer who was a director and producer of the Afghan war documentary “Restrepo,” was killed in the besieged city of Misurata, Libya, on Wednesday, and three photographers working beside him were wounded, one fatally, when they came under fire at the city’s front lines.

Chris Hondros of the Getty Images photo agency died later of devastating brain trauma. Guy Martin, who was filing photographs to the Panos agency, suffered a severe pelvic wound, said Andre Liohn, a colleague who was at the triage center where the photographers were rushed by rebels after they were struck.

Mr. Hondros suffered an extensive loss of brain tissue and was revived twice before being triaged from care. He spent several hours in a coma and died after 10 p.m., Mr. Liohn said.

Mr. Martin, a British citizen, underwent vascular surgery Wednesday night, according to the same account. As the night progressed, Mr. Liohn said that Mr. Martin’s bleeding had been stopped and that his prospects had improved, though a doctor said his condition was not yet stable.