Geographic map of the fossil location South of Erfoud, between the villages of Taouz and Begaa, Errachidia Province, Morocco; the main localities and landscape elements cited in the text are shown in the map. The asterisk marks the site from where the specimen MSNM V6894 was collected (modified from Ibrahim et al., 2016 ).

Materials and Methods

The specimen described herein was legally collected and transported to Italy together with the material published by Alessandrello & Bracchi (2003), in agreement with the Geological Service of Morocco, and permanently deposited in the Vertebrate Palaeontological Collection of the Museo di Storia Naturale di Milano (MSNM V), where it is catalogued as MSNM V6894. In conformity with Weishampel, Dodson & Osmólska (2004), we adopt the following anatomical terms of the Nomina Anatomica Veterinaria (NAV 1994) and the Nomina Anatomica Avium (NAA 1993): plantar (opposite to the back), dorsal (toward the back), proximal (toward the mass of the body), and distal (away from the mass of the body). Specimen images were taken with a Canon PowerShot S50, mounted on an ocular tube attached to a Leica MS5 stereomicroscope with Plan Apo 1.0× objective and carrier AX, then stacked with software Combine ZP.

Systematic Palaeontology

DINOSAURIA Owen, 1842

THEROPODA Marsh, 1881

SPINOSAURIDAE Stromer, 1915

SPINOSAURUS Stromer, 1915

cf. Spinosaurus aegyptiacus Stromer, 1915

MSNM V6894 strongly resembles the pedal ungual phalanges associated to diagnosable skeletal remains of specimen FSAC-KK18888, described by Ibrahim et al. (2014) and defined as the neotype of S. aegyptiacus. MSNM V6894 shares with FSAC-KK18888 the following diagnostic characters: pedal unguals with flat plantar surface; pedal unguals broader than deep with length almost four times of the proximal depth. The overall morphology, proportions, and pattern of furrows are also very similar (see “Description and comparisons”). Following Ibrahim et al. (2014) we refer the ungual MSNM V6894 to cf. S. aegyptiacus. The variability found in cervicodorsal vertebrae (Evers et al., 2015) and quadrates (Hendrickx, Mateus & Buffetaut, 2016) might indicate a higher diversity among the spinosaurid material from the Albian–Cenomanian of North Africa than previously recognized. This proportional and morphological diversity may be related to individual variability or sexual dimorphism, or it could be above the species level. However, taking into account the low number of the known specimens, their low degree of completeness, their apparently strict taxonomic affinities, their occurrence in the same strata (or, more often, their uncertain stratigraphic provenance), and all the difficulties and controversies in investigating these aspects and, ultimately, in defining a species in palaeontology, we prefer to regard all the spinosaurid material (including pedal unguals) from the Kem Kem Beds as belonging to cf. S. aegyptiacus, pending more complete, articulated remains and reliable geological data. Further comments on this topic are beyond the purpose of this paper.

Locality: MSNM V6894 comes from some kilometers south of Erfoud, between the villages of Taouz and Begaa, Errachidia Province, Morocco (Fig. 1).

Horizon: Kem Kem Beds, Cenomanian, Upper Cretaceous (Sereno et al., 1996). The specimen was collected by G. Pasini (2008, personal communication) together with rostral teeth of the Aptian–Cenomanian elasmobranch Onchopristis sp. (Rage & Cappetta, 2002; Russell, 1996).