Downing Street has responded with fury to MPs’ refusal to allow Boris Johnson to shut parliament down for a second time just days after it was reopened by his humiliating Supreme Court defeat.

The House of Commons‘ rejection – by a margin of 306 to 289 – of Mr Johnson’s request for a six-day recess lasting until Thursday has thrown plans for next week’s Conservative Party conference into disarray.

Conservative sources insisted that the four-day gathering in Manchester – opening on Sunday and due to culminate in Mr Johnson’s first address to delegates as leader on Wednesday – will go ahead as planned.

But a senior No 10 source blasted as “cynical” the MPs who voted to block the recess, including some former Tories expelled for their rebellion over Brexit.

“Conference is a part of our democratic process and it’s a massive moment for our members,” said the source. “The fact that a group of political parties have got together to try to undermine this for another political party doesn’t reflect well on them.

“The public will see a broken, zombie parliament determined to block everything, whether it is Brexit or the party conference.”

Supreme Court verdict: Boris Johnson prorogation unlawful Show all 16 1 /16 Supreme Court verdict: Boris Johnson prorogation unlawful Supreme Court verdict: Boris Johnson prorogation unlawful The Supreme Court has ruled against Boris Johnson by declaring his government unlawfully shut down parliament for five weeks EPA Supreme Court verdict: Boris Johnson prorogation unlawful Lady Hale handed down the historic verdict, hailed by opposition MPs and anti-Brexit campaigners. She said all 11 judges were unanimous in deciding that the case is “justiciable”, so the government loses that part of the argument. “The court is bound to conclude therefore that the decision to advise Her Majesty to prorogue was unlawful because it had the effect of frustrating or preventing the ability of Parliament to carry out its constitutional functions.” EPA/Supreme Court Supreme Court verdict: Boris Johnson prorogation unlawful Gina Miller and her team react outside the Supreme Court are the verdict. She said: “Today is not a win for any individual or cause. It is a win for Parliamentary sovereignty, the separation of powers and independence of our British courts. Crucially, today’s ruling confirms that we are a nation governed by the rule of law, laws that everyone, even the Prime Minister, are subject to." PA Supreme Court verdict: Boris Johnson prorogation unlawful A person dressed as a caricature of British Prime Minister Boris Johnson in a prison uniform stands outside the Supreme Court AP Supreme Court verdict: Boris Johnson prorogation unlawful Labour leader Jeremy Corbyn called for Mr Johnson to “consider his position” following the landmark decision, while Lib Dem leader Jo Swinson said he was “not fit to be prime minister” Getty Supreme Court verdict: Boris Johnson prorogation unlawful Scottish National Party (SNP) Westminster leader Ian Blackford raises his arms as he comes outside. He said: "This is an absolutely stunning judgement by the Supreme Court today." He went on to say, "we all want to get back to work, and quite frankly, on the back of this, Boris Johnson must resign immediately." AFP/Getty Supreme Court verdict: Boris Johnson prorogation unlawful Crowds outside celebrated the verdict AP Supreme Court verdict: Boris Johnson prorogation unlawful Delegates at the Labour party conference applaud after hearing the news AFP/Getty Supreme Court verdict: Boris Johnson prorogation unlawful Speaker John Bercow said MPs must now “convene without delay” and confirmed the Commons would return at 11.30am on Wednesday AFP/Getty Supreme Court verdict: Boris Johnson prorogation unlawful Gina Miller said: "“As a result of this judgment, Parliament is open, it was never prorogued. I urge MPs to get back to work immediately.” AP Supreme Court verdict: Boris Johnson prorogation unlawful MPs Anna Soubry, Liz Saville Roberts and Caroline Lucas, together with SNP leader Ian Blackford, react. Green MP Caroline Lucas has said the Supreme Court’s decision is “just the start” Reuters Supreme Court verdict: Boris Johnson prorogation unlawful “The UK deserves a Prime Minister and a Government who act with honesty, integrity and in a manner consistent with our constitution, at all times." AFP/Getty Supreme Court verdict: Boris Johnson prorogation unlawful Anti-Brexit supporters react as they gather outside the Supreme Court in London, Tuesday, Sept. 24, 2019 as it makes it's decision on the legality of Prime Minister Boris Johnson's five-week suspension of Parliament. (AP Photo/Frank Augstein) Frank Augstein AP Supreme Court verdict: Boris Johnson prorogation unlawful The SNP’s Joanna Cherry QC says: “Boris Johnson’s position is untenable and he should have the guts to resign.” PA Supreme Court verdict: Boris Johnson prorogation unlawful epa07865617 Protesters react outside the Supreme Court after the result of a hearing on the prorogation of parliament, in London, Britain, 19 September 2019. The Supreme Court ruled that the suspension of parliament by British Prime Minister Boris Johnson was not lawful. EPA/NEIL HALL NEIL HALL EPA Supreme Court verdict: Boris Johnson prorogation unlawful Tom Tugendhat MP returned to the chamber in the House of Commons within minutes of the Supreme Court ruling PA

The government’s recess motion was tabled just two days after the Supreme Court forced the reconvening of the Commons by ruling that Mr Johnson’s unprecedented five-week prorogation of parliament was unlawful.

And the No 10 response leaves no doubt that Mr Johnson regards MPs’s refusal to close Westminster’s doors once again as an act of revenge for the earlier closure.

His apparent determination to deliver his leader’s speech to the faithful in Manchester means that he will miss what would have been only his second session of PMQs since taking office two months ago, and will be regarded by many MPs as a snub to parliament.

He is likely to send a minister – probably first secretary of state Dominic Raab – to answer MPs’ questions in his place.

Recess differs from prorogation in that parliamentary business such as the tabling of questions can continue during MPs’ absence from the House, and bills which have not completed their passage through parliament do not fail.

Its denial to the Tories raises the prospect of MPs having to shuttle by train between London and Manchester to fulfil their party and parliamentary duties.

Immediately after the defeat, Leader of the House Jacob Rees-Mogg announced Commons business for next week, which included the second reading on Wednesday of the Domestic Abuse Bill, which had been dropped as a result of Mr Johnson’s abortive prorogation.

Relatively uncontroversial measures were listed for Monday and Tuesday, in an apparent effort to minimise disruption to the Conservative conference.

Mr Rees-Mogg also issued an offer to opposition parties to allow them Commons time on any day of the coming week to table a vote of no confidence in the government and trigger an election – something they are refusing to do because of the fear it would allow Mr Johnson to crash the UK out of the EU without a deal on 31 October.

It appears that the 17-vote defeat was increased by at least one MP opposing the government because of anger over Mr Johnson’s approach in Wednesday’s fiery debate over the court ruling.

Independent MP Nick Boles, who quit the Tories over Brexit, said: “I had planned to abstain on today’s motion for a short recess during the Conservative Party conference. I’m not sure if it’s the effect he intended but after the PM’s behaviour yesterday I think it is crucial that he be held to account at PMQs next Wednesday. So I will vote against.”

Mr Johnson’s official spokesman said: “The prime minister is disappointed that MPs have taken this decision. For many years it has been the case that parliament has been in recess so that parties can hold their party conferences.”