LONDON (Reuters) - Britain’s Electoral Commission is investigating whether a leading anti-EU campaigner breached referendum finance rules, after speculation mounted that Russia may have meddled in the Brexit vote.

FILE PHOTO: British businessman Arron Banks, who has funded the Leave.EU campaign, is seen during the opening day of the United Kingdom Independence Party (UKIP) annual conference at Doncaster Racecourse in Doncaster, northern Britain September 25, 2015. REUTERS/Andrew Yates/File Photo

Arron Banks, a major donor to the anti-EU campaign who was pictured with Donald Trump and leading Brexiteer Nigel Farage outside a gilded elevator soon after Trump’s 2016 U.S. presidential election victory, denied the allegations.

The Electoral Commission, which is already looking at whether Banks’ pro-Brexit Leave.EU group received any impermissible donations, said its new investigation would examine whether he was the true source of loans to a campaigner.

The investigation cannot overturn the referendum, though Banks, a 51-year-old insurance tycoon, said it was an attempt by the “Remain establishment” to discredit the Brexit result.

“Allegations of Brexit being funded by the Russians ... are complete bollocks [rubbish] from beginning to end,” Banks said in an emailed statement, signing off “nostrovia”, a version of “na zdorovye”, Russian for “cheers!”.

When asked about the investigation, Prime Minister Theresa May told parliament: “We take very seriously issues of Russian intervention, or Russian attempts to intervene, in electoral processes or in the democratic processes of any country.”

The announcement of the investigation comes after Ben Bradshaw, a lawmaker from the opposition Labour Party, asked the government to look into reports by advocacy group Open Democracy that the origin of some campaign funds was unclear.

Bradshaw said he was also concerned about what he said were significant British connections in the U.S. investigation into whether the Trump campaign colluded with Russian efforts to meddle in the presidential election.

Russia denies meddling in Brexit or the U.S. election and Trump denies any collusion with Russia.

In the June 23, 2016 referendum, 17.4 million votes, or 51.9 percent of votes cast, backed leaving the EU while 16.1 million votes, or 48.1 percent of votes cast, backed staying, a result that defied opinion polls.

VOTER CONFIDENCE

The Electoral Commission, which did not mention Russia, said it was looking at whether a company called Better for the Country Limited (BFTCL) - of which Banks was a director - was the true source of donations made to campaigners.

BFTCL was not registered as a permitted participant in the referendum but five registered campaigning groups reported receiving donations from it totalling 2.4 million pounds, the Commission said.

Banks, who was a registered permitted participant, gave three non-commercial loans to Leave.EU, totalling 6 million pounds.

Participants in the referendum were only allowed to accept donations from donors that conformed to a strict set of rules, for example they could not be based outside the UK.

The Commission said in April it was investigating Leave.EU’s funding as well as looking at whether its spending return was complete.

“Questions over the legitimacy of funding provided to campaigners at the referendum risks causing harm to voters’ confidence,” said Bob Posner, director of political finance and regulation at the Commission.

“It is therefore in the public interest that the Electoral Commission seeks to ascertain whether or not impermissible donations were given to referendum campaigners and if any other related offences have taken place,” he said.

The Commission has the power to impose fines and other sanctions if it finds rules were broken.