1. Krause, D. W., Prasad, G. V. R., von Koenigswald, W., Sahni, A. & Grine, F. E. Cosmopolitanism among Gondwanan Late Cretaceous mammals. Nature 390, 504–507 (1997).

2. Sampson, S. D. et al. Predatory dinosaur remains from Madagascar: implications for the Cretaceous biogeography of Gondwana. Science 280, 1048–1051 (1998).

3. Sereno, P. C., Wilson, J. A. & Conrad, J. L. New dinosaurs link southern landmasses in the mid-Cretaceous. Proc. R. Soc. Lond. B 217, 1325–1330 (2004).

4. Gheerbrant, E. & Rage, J.-C. Paleobiogeography of Africa: how distinct from Gondwana and Laurasia? Palaeogeogr. Palaeoclimatol. Palaeoecol. 241, 224–246 (2006).

5. Upchurch, P. Gondwanan break-up: legacies of a lost world? Trends Ecol. Evol. 23, 229–236 (2008).

6. Ali, J. R. & Krause, D. W. Late Cretaceous bioconnections between Indo-Madagascar and Antarctica: refutation of the Gunnerus Ridge causeway hypothesis. J. Biogeogr. 38, 1855–1872 (2011).

7. Ezcurra, M. D. & Agnolín, F. L. A new global palaeobiogeographical model for the late Mesozoic and early Tertiary. Syst. Biol. 61, 553–566 (2012).

8. Csiki-Sava, Z., Buffetaut, E., Ősi, A., Pereda-Suberbiola, X. & Brusatte, S. L. Island life in the Cretaceous – faunal composition, biogeography, evolution, and extinction of land-living vertebrates on the Late Cretaceous European archipelago. ZooKeys 469, 1–161 (2015).

9. Gorscak, E. & O’Connor, P. M. Time-calibrated models support congruency between Cretaceous continental rifting and titanosaurian evolutionary history. Biol. Lett. 12, 20151047 (2016).

10. Le Loeuff, J. The Campano-Maastrichtian vertebrate faunas from southern Europe and their relationships with other faunas in the world; palaeobiogeographical implications. Cretaceous Res. 12, 93–114 (1991).

11. Pereda-Suberbiola, X. Biogeographical affinities of Late Cretaceous continental tetrapods of Europe: a review. Bull. Soc. Geol. Fr. 180, 57–71 (2009).

12. Churcher, C. S. Giant Cretaceous lungfish Neoceratodus tuberculatus from a deltaic environment in the Quseir (=Baris) Formation of Kharga Oasis, Western Desert of Egypt. J. Vertebr. Paleontol. 15, 845–849 (1995).

13. Mahmoud, M. S. Palynological dating of the Quseir Formation, Kharga Oasis (Egypt). Arab. Gulf J. Sci. Res. 16, 267–281 (1998).

14. Mahmoud, M. S. Palynology and palaeoenvironment of the Quseir Formation (Campanian) from central Egypt. J. Afr. Earth Sci. 36, 135–148 (2003).

15. El Atfy, H., Sallam, H., Jasper, A. & Uhl, D. The first evidence of paleo-wildfire from the Campanian (Late Cretaceous) of North Africa. Cretaceous Res. 57, 306–310 (2016).

16. Sallam, H. M. et al. Vertebrate paleontological exploration of the Upper Cretaceous succession in the Dakhla and Kharga Oases, Western Desert, Egypt. J. Afr. Earth Sci. 117, 223–234 (2016).

17. D’Emic, M. D. The early evolution of titanosauriform sauropod dinosaurs. Zool. J. Linn. Soc. 166, 624–671 (2012).

18. Wilson, J. A. Sauropod dinosaur phylogeny: critique and cladistic analysis. Zool. J. Linn. Soc. 136, 217–276 (2002).

19. Mannion, P. D., Upchurch, P., Barnes, R. N. & Mateus, O. Osteology of the Late Jurassic Portuguese sauropod dinosaur Lusotitan atalaiensis (Macronaria) and the evolutionary history of basal titanosauriforms. Zool. J. Linn. Soc. 168, 98–206 (2013).

20. Curry Rogers, K. in The Sauropods: Evolution and Paleobiology (eds Curry Rogers, K. & Wilson, J. A.) 50–103 (Univ. California Press, Berkeley, 2005).

21. Kennedy, W. J., Klinger, H. C. & Mateer, N. J. First record of an Upper Cretaceous sauropod dinosaur from Zululand, South Africa. S. Afr. J. Sci. 83, 173–174 (1987).

22. Rauhut, O. W. M. & Werner, C. First record of a Maastrichtian sauropod dinosaur from Egypt. Palaeontol. Afr. 34, 63–67 (1997).

23. Mateus, O. et al. Angolatitan adamastor, a new sauropod dinosaur and the first record from Angola. An. Acad. Bras. Cienc. 83, 221–233 (2011).

24. Pereda Suberbiola, X., Bardet, N., Iarochène, M., Bouya, B. & Amaghzaz, M. The first record of a sauropod dinosaur from the Late Cretaceous phosphates of Morocco. J. Afr. Earth Sci. 40, 81–88 (2004).

25. Curry Rogers, K. & Forster, C. A. The skull of Rapetosaurus krausei (Sauropoda: Titanosauria) from the Late Cretaceous of Madagascar. J. Vertebr. Paleontol. 24, 121–144 (2004).

26. Le Loeuff, J. in Thunder-Lizards: The Sauropodomorph Dinosaurs (eds Tidwell, V. & Carpenter, K.) 115–137 (Indiana Univ. Press, Bloomington, 2005).

27. Wilson, J. A. Redescription of the Mongolian sauropod Nemegtosaurus mongolienesis Nowinski (Dinosauria: Saurischia) and comments on Late Cretaceous sauropod diversity. J. Syst. Palaeontol. 3, 283–318 (2005).

28. Martínez, R. D. F. et al. A basal lithostrotian titanosaur (Dinosauria: Sauropoda) with a complete skull: implications for the evolution and paleobiology of Titanosauria. PLoS. ONE 11, e0151661 (2016).

29. Wilson, J. A., Pol, D., Carvalho, A. B. & Zaher, H. The skull of the titanosaur Tapuiasaurus macedoi (Dinosauria: Sauropoda), a basal titanosaur from the Lower Cretaceous of Brazil. Zool. J. Linn. Soc. 178, 611–662 (2016).

30. Machado, E. B., Avilla, L. S., Nava, W. R., Campos, D. A. & Kellner, A. W. A. A new titanosaur sauropod from the Late Cretaceous of Brazil. Zootaxa 3701, 301–321 (2013).

31. Borsuk-Bialynicka, M. A new camarasaurid sauropod Opisthocoelicaudia skarzynskii gen. n., sp. n. from the Upper Cretaceous of Mongolia. Palaeontol. Pol. 37, 5–63 (1977).

32. Díez Díaz, V., Pereda Suberbiola, X. & Sanz, J. L. Appendicular skeleton and dermal armour of the Late Cretaceous titanosaur Lirainosaurus astibiae (Dinosauria: Sauropoda) from Spain. Palaeontol. Electron. 16, 19A (2013).

33. Gorscak, E., O’Connor, P. M., Stevens, N. J. & Roberts, E. M. The basal titanosaurian Rukwatitan bisepultus (Dinosauria, Sauropoda) from the middle Cretaceous Galula Formation, Rukwa Rift Basin, southwestern Tanzania. J. Vertebr. Paleontol. 34, 1133–1154 (2014).

34. González Riga, B. J., Lamanna, M. C., Ortiz David, L. D., Calvo, J. O. & Coria, J. P. A gigantic new dinosaur from Argentina and the evolution of the sauropod hind foot. Sci. Rep. 6, 19165 (2016).

35. Torsvik, T. H., Rousse, S., Labails, C. & Smethurst, M. A. A new scheme for the opening of the South Atlantic Ocean and the dissection of an Aptian salt basin. Geophys. J. Int. 177, 1315–1333 (2009).

36. Seton, M. et al. Global continental and ocean basin reconstructions since 200 Ma. Earth-Sci. Rev. 113, 212–270 (2012).

37. Jacobs, L. L., Winkler, D. A. & Gomani, E. M. Cretaceous dinosaurs of Africa: examples from Cameroon and Malawi. Mem. Qld Mus. 39, 595–610 (1996).

38. O’Connor, P. M. et al. A new vertebrate fauna from the Cretaceous Red Sandstone Group, Rukwa Rift Basin, southwestern Tanzania. J. Afr. Earth Sci. 44, 277–288 (2006).

39. Moody, R. T. J., & Sutcliffe, P. J. C. The Cretaceous deposits of the Iullemmeden Basin of Niger, central West Africa. Cretaceous Res. 12, 137–157 (1991).

40. Lamanna, M. C., Smith, J. B., Attia, Y. & Dodson, P. From dinosaurs to dyrosaurids (Crocodyliformes): removal of the post-Cenomanian (Late Cretaceous) record of Ornithischia from Africa. J. Vertebr. Paleontol. 24, 764–768 (2004).

41. O’Connor, P. M., Sertich, J. J. W. & Manthi, F. K. A pterodactyloid pterosaur from the Upper Cretaceous Lapurr sandstone, West Turkana, Kenya. An. Acad. Bras. Cienc. 83, 309–315 (2011).

42. Le Loeuff, J. European titanosaurids. Rev. Paléobiol. 7, 105–117 (1993).

43. Rabi, M., Tong, H. & Botfalavi, G. A new species of the side-necked turtle Foxemys (Pelomedusoides: Bothremydidae) from the Late Cretaceous of Hungary and the historical biogeography of the Bothremydini. Geol. Mag. 149, 662–674 (2012).

44. Miguel, R., Gallo, V. & Morrone, J. J. Distributional patterns of †Mawsoniidae (Sarcopterygii: Actinistia). An. Acad. Bras. Cienc. 86, 159–170 (2014).

45. Tortosa, T. et al. A new abelisaurid dinosaur from the Late Cretaceous of southern France: palaeobiogeographical implications. Ann. Paleontol. 100, 63–86 (2014).

46. Sanz, J. L., Powell, J. E., Le Loeuff, J., Martínez, R. & Pereda-Suberbiola, X. Sauropod remains from the Upper Cretaceous of Laño (northcentral Spain). Titanosaur phylogenetic relationships. Estud. Mus. Cienc. Nat. Alava 14, 235–255 (1999).

47. Díez Díaz, V. et al A new titanosaur (Dinosauria, Sauropoda) from the Upper Cretaceous of Lo Hueco (Cuenca, Spain). Cretaceous Res. 68, 49–60 (2016).

48. Lewis, P. O. A likelihood approach to estimating phylogeny from discrete morphological character data. Syst. Biol. 50, 913–925 (2001).

49. Matzke, N. J. Probabilistic historical biogeography: new models for founder-event speciation, imperfect detection, and fossils allow improved accuracy and model-testing. Front. Biogeogr. 5, 242–248 (2013).

50. Matzke, N. J. Model selection in historical biogeography reveals that founder-event speciation is a crucial process in island clades. Syst. Biol. 63, 951–970 (2014).

51. Stadler, T., Kühnert, D., Bonhoeffer, S. & Drummond, A. J. Birth–death skyline plot reveals temporal changes of epidemic spread in HIV and hepatitis C virus (HCV). Proc. Natl. Acad. Sci. USA 110, 228–233 (2013).

52. Blakey, R. SIA3584: Paleogeographic Earth Reconstruction 90 Million Years Ago (2010, accessed 10 January 2018); http://scienceviews.com/photo/library/SIA3584.html