That is alot of stellar detail you put into this. I do however have a suggestion that may help it pop a bit. I see in some areas you do have some darker lineart. Perhaps if you did the dragon's left horn with a darker outline, fading into his head. and then with his left wing, have a darker line go along his shoulders and fade it into where it meets the body. That I think will make it pop out of the paper. Well my opinion anyway. but its all good as is.