Behaviour category – highly commended. Dolphins Hunting by Greg Lecoeur (France). Location: Port Saint Johns, South Africa Sardines, a crucial part of the food chain for many marine animals, are in sharp decline. During their migration along the coast, many predators work together to hunt them but, Lecoeur says, “the action is more and more unpredictable”. He spent several days on the ocean to have just one chance and got this shot. As the judge put it: “Dolphins with sardines spilling out of their mouths. What more can you ask for?”

Photograph: Greg Lecoeur/UPY2017