The NFL has disciplined 33 players from the Pittsburgh Steelers and Cleveland Browns -- with fines totaling $732,422 -- as penalty for the brawl that broke out in the closing seconds of their game on Nov. 14.

The league on Saturday fined 29 players -- including Pittsburgh's Minkah Fitzpatrick and Cleveland's Jarvis Landry -- $3,507 apiece for "entering the fight area."

Browns defensive end Myles Garrett was suspended indefinitely and fined $45,623 for ripping off and swinging Mason Rudolph's helmet, hitting the Steelers quarterback in the head with it during Cleveland's 21-7 victory.

Disciplined Team Fine Cleveland Browns $250,000 Pittsburgh Steelers $250,000 Tyson Alualu PIT $3,507 Mark Barron PIT $3,507 Tony Brooks-James PIT $3,507 Juston Burris CLE $3,507 T.J. Carrie CLE $3,507 Bud Dupree PIT $3,507 Terrell Edmunds PIT $3,507 Trey Edmunds PIT $3,507 Matt Feiler PIT $3,507 Minkah Fitzpatrick PIT $3,507 Ramon Foster PIT $3,507 Myles Garrett* CLE $45,623 Mike Hilton PIT $3,507 Johnny Holton PIT $3,507 Tevin Jones PIT $3,507 Kameron Kelly PIT $3,507 Jarvis Landry CLE $3,507 Devaroe Lawrence CLE $3,507 Larry Ogunjobi* CLE Rescinded Maurkice Pouncey* PIT $35,096 Sheldrick Redwine CLE $3,507 Mason Rudolph PIT $50,000 Jaylen Samuels PIT $3,507 Chris Smith CLE $3,507 Robert Spillane PIT $3,507 Cameron Sutton PIT $3,507 Chad Thomas CLE $3,507 Denzel Ward CLE $3,507 James Washington PIT $3,507 T.J. Watt PIT $3,507 Greedy Williams CLE $3,507 Vince Williams PIT $3,507 Mack Wilson CLE $3,507 *Suspended

Garrett lost his appeal this week and stands to lose about $1.2 million in salary for missing the six remaining games.

Rudolph, who was not suspended, was slapped with the biggest fine, of $50,000. Pittsburgh center Maurkice Pouncey, who won an appeal reducing his suspension from three to two games, was fined $35,096 for fighting.

Browns defensive tackle Larry Ogunjobi was suspended for one game, but his $10,527 fine was rescinded upon appeal.

In addition, both the Browns and Steelers franchises were fined $250,000 each.

Fines collected go to programs for former NFL players.

Three Cleveland players were also fined for plays that happened before the melee.

Safety Damarious Randall was fined $28,075 for unnecessary roughness and was ejected for a hit to the head of Steelers receiver Diontae Johnson, which knocked Johnson out of the game with a concussion.

Defensive end Chad Thomas was fined $21,056 for a hit on Rudolph. From the same play, cornerback Greedy Williams was fined $28,075 for unnecessary roughness on a hit that left Steelers receiver JuJu Smith-Schuster with a concussion as well.

The Browns travel to Pittsburgh for the rematch Dec. 1.