General features All rodents possess constantly growing rootless incisors that have a hard enamel layer on the front of each tooth and softer dentine behind. The differential wear from gnawing creates perpetually sharp chisel edges. Rodents’ absence of other incisors and canine teeth results in a gap, or diastema, between incisors and cheek teeth, which number from 22 (5 on each side of the upper and lower jaws) to 4, may be rooted or rootless and ever-growing, and may be low- or high-crowned. The nature of the jaw articulation ensures that incisors do not meet when food is chewed and that upper and lower cheek teeth (premolars and molars) do not make contact while the animal gnaws. Powerful and intricately divided masseter muscles, attached to jaw and skull in different arrangements, provide most of the power for chewing and gnawing. capybara Capybara (Hydrochoerus hydrochaeris). © Wilfredo Rodríguez (A Britannica Publishing Partner) The range in body size between the mouse (18 grams [0.64 ounce], body 12 cm [4.7 inches] long) and the marmot (3,000 grams, body 50 cm long) spans the majority of living rodents, but the extremes are remarkable. One of the smallest is Delany’s swamp mouse (Delanymys brooksi), associated with bamboo in the marshes and mountain forests in Africa. It weighs 5 to 7 grams, and the body is 5 to 6 cm long. The largest is the capybara (Hydrochoerus hydrochaeris) of Central and South America, which weighs 35 to 66 kg (77 to 146 pounds) and stands 50 to 60 cm at the shoulder, with a body 100 to 135 cm long. Some extinct species were even larger, attaining the size of a black bear or small rhinoceros. The largest rodent ever recorded, Josephoartigasia monesi, lived some two to four million years ago, during the Pleistocene and Pliocene epochs; by some estimates it grew to a length of about 3 metres (10 feet) and weighed nearly 1,000 kg.

Natural history Rodents may be diurnal, nocturnal, or sometimes active part of the day and night. Although some species are herbivorous, diets of most include vegetable and animal matter. Others are opportunistic generalists, and some are specialized predators, not only of arthropods (see grasshopper mouse) but sometimes of vertebrates. Food is either eaten where gathered or carried to burrows and stored (see pocket gopher, pocket mouse, African pouched rat, and hamster). Species living in arid habitats and on oceanic islands are able to obtain their water requirements from their food. A wide variety of shelters are used or constructed; these range from tree holes, rock crevices, or simple burrows to hidden nests on the forest floor, leaf and stick structures in tree crowns, mounds of cut vegetation built in aquatic environments, or complex networks of tunnels and galleries. Rodents may be active all year or enter periods of dormancy or deep hibernation. Breeding time and frequency, length of gestation, and litter size vary widely, but two of the most prolific are both associated with humans. The brown rat (Rattus norvegicus) can give birth to litters of up to 22 offspring, and the house mouse (Mus musculus) can produce up to 14 litters annually. Population size may remain stable or fluctuate, and some species, most notably lemmings, migrate when populations become excessively large.