He is called the “father of the American cavalry,” a Polish-born Revolutionary War hero who fought for American independence under George Washington and whose legend inspired the dedication of parades, schools, roads and bridges.

But for more than 200 years, a mystery persisted about his final resting place. Historical accounts suggested the cavalryman, Casimir Pulaski, had been buried at sea, but others maintained he was buried in an unmarked grave in Savannah, Ga.

Researchers believe they have found the answer — after coming to another significant discovery: The famed general was most likely intersex.

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New evidence suggests that although Pulaski identified and lived as a man, biologically, he did not fit into the binary definitions of male and female, a twist that helps explain why scientists could not previously identify his remains. The revelatory findings are detailed in a new documentary, “The General Was Female?,” which aired on the Smithsonian Channel on April 8.