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Wales centre Jonathan Davies will miss the rest of the autumn campaign after suffering a big injury blow in the first half of the 24 - 15 defeat against South Africa.

Davies was taken off after just 12 minutes of a bruising encounter against the Springboks after suffering a damaged pectoral muscle.

Coach Warren Gatland confirmed after the match he will be unavailable for the forthcoming matches against Argentina, Tonga and Australia.

The loss of the Lions man is a huge blow, with fellow Lion Jamie Roberts also missing from the Welsh midfield.

Davies was the most influential player on the pitch in the opening exchanges, his departure leaving Wales rattled. If the Scarlets man has to have an operation on the problem, he could be out for five months.

Liam Williams and Adam Jones were also both taken off inside the opening half hour, with the latter set to be out for at least a couple of weeks with a calf strain that forced him onto crutches after the match.

Williams' availability remains unclear after he suffered a head injury in the run up to South Africa's opening score.

Gatland said: “The injuries are relatively serious. Adam Jones is a calf strain, Jonathan Davies is a pectoral muscle which will keep him out for the rest of the campaign, Liam Williams has had a bump to the head and will be assessed. Scott Andrews is OK.

“Absolutely. He was a big loss to us in the first half, he had made a couple of breaks and been strong on his feet.

“He is an important player for us and he gives us that left-foot kicking option.

“He is very disappointed. We’ve spoken to the physio and he said it could settle by the back end of December, but if he needs an operation it could be five months and mean he misses the Six Nations as well."