“It’s hard to imagine losing something longer than a pickup truck,” he added, referring to what he called Berkeley’s “amazing incompetence.”

“It’s astounding,” he said.

In correspondence with the federal government, Andrew Goldblatt, who has the stressful-sounding title of assistant risk manager for the university, described the sale of the Johnson piece as “an error of ignorance.”

“We do regret it,” Mr. Goldblatt said in an interview. “Something went wrong, and it just cascaded.”

Johnson (1888-1967) is considered one of the finest sculptors of the Harlem Renaissance, though he spent most of his life in the Bay Area. He was never able to earn a living purely from his art, but in recent years interest in him has resurged, said Gwendolyn DuBois Shaw, an associate professor of American art at the University of Pennsylvania, who is writing a book on him. In 1937, under the auspices of the W.P.A., Johnson designed two large Art Deco redwood reliefs, one of which depicted an idealized natural world of gilded gazelles, open-beaked birds, spiky-leafed plants and a boy clapping cymbals.

Designed to cover organ pipes at the old California School for the Deaf and Blind in Berkeley, this natural-world relief was affixed to a wall until 1980, when the school moved. As squatters (and rats) took shelter there, the university, which had taken over the premises, moved any valuable property to a secure basement warehouse, and the organ relief was disassembled. But one of the organ screens was misidentified as belonging to Berkeley’s graduate schools, so when the university reopened the building three years later, only one of the two Johnson reliefs was returned to its rightful place. The other remained in storage until 2009, when the university emptied the storage space in preparation for the sale of the building and transferred the relief to the university’s surplus store.