Portrait category - winner. A sand tiger shark surrounded by tiny bait fish by Tanya Houppermans (US). Location: Wreck of Caribsea, North Carolina Houppermans caught this dramatic shot while diving to the wreck of the Caribsea and seeing the fierce-looking, but docile, sand tiger sharks that frequent the wreck. On this particular day millions of tiny fish, collectively known as “bait fish”, were grouped together in an enormous bait ball above the wreck. As she moved with the shark through the water the bait fish parted way, giving her a clear shot of the underside of this beautiful shark.

Photograph: Tanya Houppermans/UPY 2018