ALEXANDRIA, Va. (Reuters) - President Donald Trump’s former campaign manager Paul Manafort pleaded not guilty on Thursday to additional criminal charges ranging from bank fraud to filing false tax returns.

U.S. Special Counsel Robert Mueller brought them as part of a wide-ranging probe into Russian interference in the 2016 election, possible obstruction of justice, and alleged financial crimes by Manafort and others.

Trump has denied that his campaign colluded with Russia.