Now, candidates must focus on outlining what people, businesses and governments can do, and how their administration might create incentives to promote resiliency. Mr. Ulvert said that word — resiliency — is important for candidates looking to expand their appeal beyond liberal voters to moderates, because it eases less-engaged voters into a conversation about the environment even when that is not their top concern.

“You’re coming into Florida, into Miami-Dade, where there are persuadable voters,” he said. “The issue of resiliency is an opportunity of how you can transcend your message beyond partisan politics. Talk about it in a way where you’re bringing people to the table.”

Thinking past the primary might be impossible for the Democratic contenders now. But the eventual nominee will have to run in Florida, an important presidential battleground state, at a time when more Republicans are campaigning on the environment.

The new Republican governor, Ron DeSantis, has appointed a University of Florida biologist as the state’s first chief science officer and is hiring a chief resilience officer, whose job description includes preparing Florida “for the environmental, physical and economic impacts of climate change, especially sea-level rise.”

Mr. DeSantis campaigned not on fighting climate change but on improving water quality, including addressing pollution that contributed to toxic algae blooms. That showed voters like Jim Cass, a commercial fisherman in Key West, that Mr. DeSantis understood how human activity can harm the economy.

“That’s encouraging to me, though it’s 20 years overdue,” said Mr. Cass, 75, a Republican who voted for Mr. Trump but said he has been disappointed by his antics and by the Republican Party’s inaction in Congress.

Mr. Cass attributes changes in fish behavior directly to global warming. Over the last few years, yellowtail snapper have been spawning earlier because the water warms sooner, and king fish have stayed closer to Naples, Fla., because the waters there have gotten warm enough that they do not need to migrate farther south.