1. Woodward, A. S. A new cave man from Rhodesia, South Africa. Nature 108, 371–372 (1921).

2. Stringer, C. B. Some further notes on the morphology and dating of the Petralona hominid. J. Hum. Evol. 12, 731–742 (1983).

3. Stringer, C. The status of Homo heidelbergensis (Schoetensack 1908). Evol. Anthropol. 21, 101–107 (2012).

4. Rightmire, G. P. in Human Paleontology and Prehistory 145–159 (Springer, 2017).

5. Millard, A. R. A critique of the chronometric evidence for hominid fossils: I. Africa and the Near East 500–50 ka. J. Hum. Evol. 54, 848–874 (2008).

6. Klein, R. G. The Human Career: Human Biological and Cultural Origins (Univ. Chicago Press, 2009).

7. White, T. D. et al. Pleistocene Homo sapiens from Middle Awash, Ethiopia. Nature 423, 742–747 (2003).

8. Richter, D. et al. The age of the hominin fossils from Jebel Irhoud, Morocco, and the origins of the Middle Stone Age. Nature 546, 293–296 (2017).

9. Hublin, J.-J. et al. New fossils from Jebel Irhoud, Morocco and the pan-African origin of Homo sapiens. Nature 546, 289–292 (2017).

10. Berger, L. R., Hawks, J., Dirks, P. H., Elliott, M. & Roberts, E. M. Homo naledi and Pleistocene hominin evolution in subequatorial Africa. eLife 6, e24234 (2017).

11. Dirks, P. H. G. M. et al. The age of Homo naledi and associated sediments in the Rising Star Cave, South Africa. eLife 6, e24231 (2017).

12. Galway-Witham, J. et al. Aspects of human physical and behavioural evolution during the last 1 million years. J. Quat. Sci. 34, 355–378 (2019).

13. Stringer, C. B. The origin and evolution of Homo sapiens. Phil. Trans. R. Soc. B 371 2015.0237 (2016).

14. Lacruz, R. S. et al. The evolutionary history of the human face. Nat. Ecol. Evol. 3, 726–736 (2019).

15. White, F. Notes on a cave containing fossilized bones of animals, worked pieces of bone, stone implements, and quartzite pebbles, found in a kopje or small hill composed of zinc and lead ores, at Broken Hill, North-Western Rhodesia. Proc. Rhod. Sci. Assoc. 7, 13–21 (1908).

16. Oakley, K. P., Campbell, B. G. & Molleson, T. I. Catalogue of Fossil Hominids, Part I: Africa (Natural History Museum Publications, 1977).

17. Trinkaus, E. The human tibia from Broken Hill, Kabwe, Zambia. Paleoanthropology 2009, 145–165 (2009).

18. Stringer, C. B. An archaic character in the Broken Hill innominate E. 719. Am. J. Phys. Anthropol. 71, 115–120 (1986).

19. Clark, J. D., Oakley, K. P., Wells, L. H. & McClelland, J. A. New studies on Rhodesian Man. J. R. Anthropol. Inst. 77, 7–32 (1947).

20. Clark, J. D. Further excavations at Broken Hill, Northern Rhodesia. J. R. Anthropol. Inst. 89, 201–232 (1959).

21. Mcbrearty, S. & Brooks, A. S. The revolution that wasn’t: a new interpretation of the origin of modern human behavior. J. Hum. Evol. 39, 453–563 (2000).

22. Barham, L., Pinto, A. & Stringer, C. Bone tools from Broken Hill (Kabwe) cave, Zambia, and their evolutionary significance. Before Farming 2, 1–16 (2002).

23. Brink, J. S. in Quaternary Environmental Change in Southern Africa: Physical and Human Dimensions (eds Knight, J. & Grab, S. W.) 284–305 (Cambridge Univ. Press, 2016).

24. Grün, R., Eggins, S., Kinsley, L., Mosely, H. & Sambridge, M. Laser ablation U-series analysis of fossil bones and teeth. Palaeogeogr. Palaeoclimatol. Palaeoecol. 416, 150–167 (2014).

25. Stringer, C. Modern human origins: progress and prospects. Phil. Trans. R. Soc. Lond. B 357, 563–579 (2002).

26. McDougall, I., Brown, F. H. & Fleagle, J. G. Stratigraphic placement and age of modern humans from Kibish, Ethiopia. Nature 433, 733–736 (2005).

27. Aubert, M. et al. Confirmation of a late middle Pleistocene age for the Omo Kibish 1 cranium by direct uranium-series dating. J. Hum. Evol. 63, 704–710 (2012).

28. Grün, R. et al. Direct dating of Florisbad hominid. Nature 382, 500–501 (1996).

29. Bräuer, G., Yokoyama, Y., Falguères, C. & Mbua, E. Modern human origins backdated. Nature 386, 337–338 (1997).

30. Scerri, E. M. L. et al. Did our species evolve in subdivided populations across Africa, and why does it matter? Trends Ecol. Evol. 33, 582–594 (2018).

31. Lorente-Galdos, B. et al. Whole-genome sequence analysis of a Pan African set of samples reveals archaic gene flow from an extinct basal population of modern humans into sub-Saharan populations. Genome Biol. 20, 77 (2019).

32. Meyer, M. et al. Nuclear DNA sequences from the Middle Pleistocene Sima de los Huesos hominins. Nature 531, 504–507 (2016).

33. Ludwig, K. R. Isoplot. Excel package version 3.76.12.02.24 http://www.bgc.org/isoplot_etc/isoplot.html (2012).

34. Harris, W. E. The finding of the Broken Hill skull: The mystery of the Great Bone Cave. London Illustrated News 679–681 (19 November 1921).

35. Hrdlička, A. The Skeletal Remains of Early Man Vol. 83 (Smithsonian Institution Miscellaneous Collections, 1930).

36. Pycraft, W. P. et al. Rhodesian Man and Associated Remains (British Museum (Natural History), 1928).

37. White, F. Letter to Smith Woodward (dated 20th November 1921) Natural History Museum Archives (1921).

38. Clark, J. D., Brothwell, D. R., Powers, R. & Oakley, K. P. Rhodesian man: notes on a new femur fragment. Man 3, 105–111 (1968).

39. Bartsiokas, A. Studies of Recent and Fossil Bone, Including X-ray Microanalysis, with Special Reference to Kabwe (Rhodesian) Man. PhD thesis, Univ. London (1989).

40. Chubb, C. E. List of vertebrate remains. Proc. Rhod. Sci. Assoc. 7, 21–23 (1908).

41. Hopwood, A. T. in Rhodesian Man and Associated Remains 70–73 (British Museum (Natural History), 1928).

42. Leakey, L. S. B. A preliminary re-assessment of the fossil fauna from Broken Hill, Northern Rhodesia, in Clark, J. Desmond, ‘Further Excavations at Broken Hill, Northern Rhodesia’. J. R. Anthropol. Inst. 89, 201–232 (1959).

43. Klein, R. G. Geological antiquity of Rhodesian man. Nature 244, 311–312 (1973).

44. Churcher, C. S. in Evolution of African Mammals (eds Maglio, V. J. & Cooke, H. B. S.) 509–535 (Harvard Univ. Press, 1978).

45. Churcher, C. S. & Richardson, M. L. in Evolution of African Mammals (eds Maglio, V. J. & Cooke, H. B. S.) 379–422 (Harvard Univ. Press, 1978).

46. Gentry, A. W. in Evolution of African Mammals (eds Maglio, V. J. & Cooke, H. B. S.) 540–572 (Harvard Univ. Press, 1978).

47. Savage, R. J. G. in Evolution of African Mammals (eds Maglio, V. J. & Cooke, H. B. S.) 249–267 (Harvard Univ. Press, 1978).

48. Lacruz, R. et al. Palaeontology and geological context of a Middle Pleistocene faunal assemblage from the Gladysvale Cave, South Africa. Palaeontol. Afr. 38, 99–114 (2002).

49. Brink, J. S. The Evolution of the Black Wildebeest (Connochaetes gnou) and Modern Large Mammal Faunas of Central Southern Africa. PhD thesis, Univ. Stellenbosch (2005).

50. Brink, J. S. et al. First hominine remains from a ~1.0 million year old bone bed at Cornelia-Uitzoek, Free State Province, South Africa. J. Hum. Evol. 63, 527–535 (2012).

51. Klein, R. G., Avery, G., Cruz-Uribe, K. & Steele, T. E. The mammalian fauna associated with an archaic hominin skullcap and later Acheulean artifacts at Elandsfontein, Western Cape Province, South Africa. J. Hum. Evol. 52, 164–186 (2007).

52. Vrba, E. S. New fossils of Alcelaphini and Caprinae (Bovidae, Mammalia) from Awash, Ethiopia, and phylogenetic analysis of Alcelaphini. Palaeontol. Afr. 34, 127–198 (1997).

53. Klein, R. G. (ed.) in Southern African Prehistory and Palaeoenvironments 107–146 (A. A. Balkema, 1984).

54. Denys, C. in African Biogeography, Climate Change and Human Evolution (eds Bromage, T. & Schrenk, F.) 226–252 (Oxford Univ. Press, 1999).

55. Mennell, F. P. & Chubb, E. C. On an African occurrence of fossil Mammalia associated with stone implements. Geol. Mag. 4, 443–448 (1907).

56. Avery, D. M. Taphonomy of micromammals from cave deposits at Kabwe (Broken Hill) and Twin Rivers in central Zambia. J. Archaeol. Sci. 29, 537–544 (2002).

57. Avery, D. M. Early and Middle Pleistocene environments and hominid biogeography; micromammalian evidence from Kabwe, Twin Rivers and Mumbwa Caves in central Zambia. Palaeogeogr. Palaeoclimatol. Palaeoecol. 189, 55–69 (2003).

58. Denys, C. in Advances in Vertebrate Paleontology “Hen to Panta” Volume in Honor of Constantin Radulescu and Petre Mihai Samson (eds Petrulescu, A. & Stiuca, E.) 75–84 (Romanian Academy, 2003).

59. Deino, A. L. & Mcbrearty, S. 40Ar/39Ar dating of the Kapthurin Formation, Baringo, Kenya. J. Hum. Evol. 42, 185–210 (2002).

60. Denys, C. & Jaeger, J. J. A biostratigraphic problem: the case of the East African Plio-Pleistocene rodent faunas. Mod. Geol. 10, 215–233 (1986).

61. Monadjem, A., Taylor, P. J., Denys, C. & Cotteril, F. Rodents of Sub-Saharan Africa: A Biogeographic and Taxonomic Synthesis (Walter de Gruyter, 2015).

62. Colangelo, P., Granjon, L., Taylor, P. J. & Corti, M. Evolutionary systematics in African gerbilline rodents of the genus Gerbilliscus: inference from mitochondrial genes. Mol. Phylogenet. Evol. 42, 797–806 (2007).

63. Dobigny, G. et al. Mitochondrial and nuclear genes-based phylogeography of Arvicanthis niloticus (Murinae) and sub-Saharan open habitats Pleistocene history. PLoS ONE 8, e77815 (2013).

64. Corti, M., Castiglia, R., Annesi, F. & Verheyen, W. Mitochondrial sequences and karyotypes reveal hidden diversity in African pouched mice (subfamily Cricetomyinae, genus Saccostomus). J. Zool. 262, 413–424 (2004).

65. McDermott, F. et al. New Late-Pleistocene uranium–thorium and ESR dates for the Singa hominid (Sudan). J. Hum. Evol. 31, 507–516 (1996).

66. Pike, A. W. G., Hedges, R. E. M. & Van Calsteren, P. U-series dating of bone using the diffusion–adsorption model. Geochim. Cosmochim. Acta 66, 4273–4286 (2002).

67. Grün, R., Yan, G., McCulloch, M. & Mortimer, G. Detailed mass spectrometric U-series analyses of two teeth from the archaeological site of Pech de l’Aze II: implications for uranium migration and dating. J. Archaeol. Sci. 26, 1301–1310 (1999).

68. Price, G. J., Feng, Y. X., Zhao, J. X. & Webb, G. E. Direct U–Th dating of vertebrate fossils with minimum sampling destruction and application to museum specimens. Quat. Geochronol. 18, 1–8 (2013).

69. Simpson, J. J. & Grün, R. Non-destructive gamma spectrometric U-series dating. Quat. Sci. Rev. 17, 1009–1022 (1998).

70. Benson, A. et al. Laser ablation depth profiling of U-series and Sr isotopes in human fossils. J. Archaeol. Sci. 40, 2991–3000 (2013).

71. Duval, M., Aubert, M., Hellstrom, J. & Grün, R. High resolution, LA-ICP-MS mapping of U and Th isotopes in an Early Pleistocene equid tooth from Fuente Nueva-3 (Orce, Andalusia, Spain). Quat. Geochronol. 6, 458–467 (2011).

72. Grün, R., Joannes-Boyau, R. & Stringer, C. Two types of CO 2 − radicals threaten the fundamentals of ESR dating of tooth enamel. Quat. Geochronol. 3, 150–172 (2008).

73. Joannes-Boyau, R. & Grün, R. A comprehensive model for CO 2 − radicals in fossil tooth enamel: implications for ESR dating. Quat. Geochronol. 6, 82–97 (2011).

74. Nathan, R. & Grün, R. Gamma dosing and shielding of a human tooth by a mandible and skull cap. Ancient TL 21, 79–84 (2003).

75. Duval, M. & Martín-Francés, L. Quantifying the impact of µCT-scanning of human fossil teeth on ESR age results. Am. J. Phys. Anthropol. 163, 205–212 (2017).

76. Grün, R. An alternative for model for open system U-series/ESR age calculations: (closed system U-series)-ESR, CSUS-ESR. Ancient TL 18, 1–4 (2000).

77. Grün, R., Schwarcz, H. P. & Chadam, J. M. ESR dating of tooth enamel: coupled correction for U-uptake and U-series disequilibrium. Nucl. Tracks Radiat. Meas. 14, 237–241 (1988).